Revision of the Australian scorpion-fly genus Harpobittacus (Mecoptera : Bittacidae)

Abstract

Harpobittacus Gerstaecker is the largest of the six genera of Australian Bittacidae. Adults occur in eastern, south-eastern and south-western Australian eucalypt woodland and coastal heathland during spring and summer and sometimes autumn. The genus contains 11 species, which are diagnosed in the present revision: H. australis (Klug) [= australis rubripes Riek, syn. nov., = corethrarius (Rambur), = intermedius (Selys-Longchamps)] (south-east Australia, including Tasmania); H. albatus Riek, stat. nov. (= limnaeus Smithers, syn. nov.) (coastal eastern Australia); H. christine, sp. nov. (inland south-east Queensland); H. tillyardi Esben-Petersen ( = nigratus Navás) (coastal eastern Australia); H. rubricatus Riek (inland south-east Australia); H. scheibeli Esben-Petersen (= brewerae Smithers, syn. nov.) (inland and coastal eastern Australia); H. septentrionis, sp. nov. (coastal north Queensland); H. nigriceps (Selys-Longchamps) (mainland south-east Australia); H. similis Esben-Petersen, H. quasisimilis, sp. nov., and H. phaeoscius Riek (all south-west Western Australia). Cladistic analysis has produced the following hypothesis of relationships: (((australis (albatus christine)) (tillyardi rubricatus)) ((similis quasisimilis) ((scheibeli septentrionis) (nigriceps phaeoscius))). Immediate sister-species show little or no overlap in their geographic distributions.