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RESEARCH ARTICLE
Contents Vol 32(2)

A new koala (Marsupialia : Phascolarctidae) from the late Oligocene Etadunna Formation, Lake Eyre Basin, South Australia

Neville S. Pledge
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South Australian Museum, Adelaide, SA 5000, Australia. Email: pledge.neville@saugov.sa.gov.au

Australian Mammalogy 32(2) 79-86 https://doi.org/10.1071/AM09014
Submitted: 12 May 2009  Accepted: 23 February 2010   Published: 26 July 2010

Abstract

An isolated upper molar represents Litokoala thurmerae sp. nov., the eighth species of phascolarctid marsupial (koalas) from the South Australian mid-Tertiary sequence, and the fourth from the late Oligocene Etadunna Formation at Lake Palankarinna. It is the smallest and oldest species, differing from L. kutjamarpensis (Stirton et al. 1967), L. kanunkaensis (Springer 1987) and L. dicktedfordi, sp. nov. (the Riversleigh specimens referred to L. kanunkaensis and L. kutjamarpensis (Black and Archer 1997; Louys et al. 2007) but described here as a new species) in size, and the almost total lack of crenulations on the surfaces of the cusps. This brings to at least five the number of probably arboreal mammal species in the Ngama Local Fauna (Pledge 1984) of Mammalon Hill, Lake Palankarinna – the others being Ektopodon stirtoni (Pledge 1986), Pildra magnus (Pledge 1987a), P. sp. cf. kutjamarpensis (ibid.), and Burramys wakefieldi (Pledge 1987b) – and further supports the riparian forest environment interpretation proposed for this part of the Etadunna sequence (Pledge 1984; Martin 2006).


Acknowledgements

I thank Professor Mike Woodburne (then University of California, Riverside) for allowing me to borrow the material of Litokoala kanunkaensis in his care, and for numerous discussions on this and other topics. Ben McHenry and Bill Aird assisted with the scanning electron microscopy. Mostly, I am grateful to my field crew, in particular Jenni Thurmer whose keen eyes spotted this and many other specimens. Thanks also to the referees for useful comments.


References

Archer, M. (1978). The nature of the molar-premolar boundary in marsupials and a reinterpretation of the homology of marsupial cheek-teeth. Memoirs of the Queensland Museum 18, 157–164.
Lee A. , and Martin R. (1988). ‘The Koala – A Natural History.’ Australian Natural History Series. (New South Wales University Press: Sydney.)

Louys, J. , Black, K. , Archer, M. , Hand, S. J. , and Godthelp, H. (2007). Descriptions of koala fossils from the Miocene of Riversleigh, northwestern Queensland and implications for Litokoala (Marsupialia, Phascolarctidae). Alcheringa 31, 99–110.
| Crossref | GoogleScholarGoogle Scholar | Luckett W. P. (1993). An ontogenetic assessment of dental homologies in therian mammals. In ‘Mammal Phylogeny: Mesozoic Differentiation, Multituberculates, Monotremes, Early Eutherians and Marsupials’. (Eds F. S. Szalay, M. J. Novacek and M. C. McKenna.) pp. 182–204. (Springer: New York.)

Martin, H. A. (2006). Cenozoic climate change and the development of the arid vegetation in Australia. Journal of Arid Environments 66, 533–563.
| Crossref | GoogleScholarGoogle Scholar | Pledge N. S. (1986). A new species of Ektopodon (Marsupialia; Phalangeroidea) from the Miocene of South Australia. In ‘Revision of the Ektopodontidae (Mammalia; Marsupialia; Phalangeroidea) of the Australian Neogene’. (Eds M. O. Woodburne and W. A. Clemens.) pp. 43–67. University of California Publications in Geological Sciences 31.

Pledge N. S. (1987 a). Pseudocheirids (Marsupialia: Pseudocheiridae) from the Middle Miocene Ngama Local Fauna of northern South Australia. In ‘Possums and Opossums: Studies in Evolution’. (Ed. M. Archer.) pp. 681–688. (Surrey Beatty & Royal Zoological Society of New South Wales: Sydney.)

Pledge N. S. (1987 b). A new species of Burramys Broom (Marsupialia: Burramyidae) from the Middle Miocene of South Australia. In ‘Possums and Opossums: Studies in Evolution’. (Ed. M. Archer.) pp. 725–728. (Surrey Beatty & Royal Zoological Society of New South Wales: Sydney.)

Springer M. S. (1987). Lower molars of Litokoala (Marsupialia: Phascolarctidae) and their bearing on phascolarctid evolution. In ‘Possums and Opossums: Studies in Evolution’. (Ed. M. Archer.) pp. 319–325. (Surrey Beatty & Royal Zoological Society of New South Wales: Sydney.)

Stirton, R. A. (1957). A new koala from the Pliocene Palankarinna fauna of South Australia. Records of the South Australian Museum 13, 71–81.
Tedford R. H. , and Woodburne M. O. (1987). The Ilariidae, a new family of vombatiform marsupials from Miocene strata of South Australia and an evaluation of the homology of molar cusps in the Diprotodontidae. In ‘Possums and Opossums: Studies in Evolution’. (Ed. M. Archer.) pp. 401–418. (Surrey Beatty & Royal Zoological Society of New South Wales: Sydney.)

Woodburne M. O. , Tedford R. H. , Archer M. , and Pledge N. S. (1987). Madakoala, a new genus and two species of Miocene koalas (Marsupialia: Phascolarctidae) from South Australia, and a new species of Perikoala. In ‘Possums and Opossums: Studies in Evolution’. (Ed. M. Archer.) pp. 293–317. (Surrey Beatty & Royal Zoological Society of New South Wales: Sydney.)

Woodburne, M. O. , MacFadden, B. J. , Case, J. A. , Springer, M. S. , Pledge, N. S. , Power, J. D. , Woodburne, J. M. , and Springer, K. B. (1993). (published 1994). Land mammal biostratigraphy and magnetostratigraphy of the Etadunna Formation (Late Oligocene) of South Australia. Journal of Vertebrate Paleontology 13, 483–515.