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Systematics, phylogeny and biogeography
RESEARCH ARTICLE

Cladistic analysis supports the monophyly of the Neotropical crab spider genus Epicadus and its senior synonymy over Tobias (Araneae : Thomisidae)

Miguel Machado A B , Renato A. Teixeira A and Arno A. Lise A
+ Author Affiliations
- Author Affiliations

A Laboratório de Aracnologia, Faculdade de Biociências, Pontifícia Universidade Católica do Rio Grande do Sul (PUCRS), Porto Alegre, Rio Grande do Sul, Brazil.

B Corresponding author. Email: machadom.arachno@gmail.com

Invertebrate Systematics 31(4) 442-455 https://doi.org/10.1071/IS16074
Submitted: 5 November 2016  Accepted: 20 February 2017   Published: 23 June 2017

Abstract

The present paper presents a cladistic analysis of the spider genus Tobias Simon, 1895. The analyses were based on a matrix with 29 terminal taxa scored for 86 morphological characters, with a dataset tested under two methodologies for character weighting (i.e. equal and implied weighting). Both analyses supported the paraphyletic relationship of Tobias with Epicadus Simon, 1895, with the former considered a junior synonym of the latter (new synonymy). Onocolus, Epicadinus and Epicadus form a clade supported by two synapomorphies, corroborating previous taxonomic assumptions regarding the proximity of these genera. Epicadus has two lineages: the ‘pustulosus clade’, comprising bark-like individuals that camouflage themselves on tree trunks and branches, and the ‘heterogaster clade’, which consists of a group of spiders that present a wide range of polychromatism and use flowers as hunting sites. Synonymies are presented and a new combination for all species previously assigned as part of Tobias is proposed. Epicadus now comprises nine species.

Additional keywords: phylogeny, Stephanopinae.


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