Free Standard AU & NZ Shipping For All Book Orders Over $80!
Register      Login
Wildlife Research Wildlife Research Society
Ecology, management and conservation in natural and modified habitats
RESEARCH ARTICLE (Open Access)

Effectiveness and costs of helicopter-based shooting of deer

Andrew J. Bengsen https://orcid.org/0000-0003-2205-4416 A , David M. Forsyth https://orcid.org/0000-0001-5356-9573 A * , Anthony Pople B , Michael Brennan B , Matt Amos B , Mal Leeson C , Tarnya E. Cox A , Bec Gray D , Ollie Orgill E , Jordan O. Hampton https://orcid.org/0000-0003-0472-3241 F H , Troy Crittle D and Kym Haebich G
+ Author Affiliations
- Author Affiliations

A Vertebrate Pest Research Unit, NSW Department of Primary Industries, 1447 Forest Road, Orange, NSW 2800, Australia.

B Biosecurity Queensland, Queensland Department of Agriculture and Fisheries, 41 Boggo Road, Dutton Park, Qld 4102, Australia.

C Local Land Services Regional Operations, 112 Market Street, Mudgee, NSW 2850, Australia.

D Biosecurity and Food Safety, NSW Department of Primary Industries, 4 Marsden Park Road, Calala, NSW 2340, Australia.

E Environment, Planning and Sustainable Development Directorate, ACT Government, Sanford Street, Mitchell, ACT 2911, Australia.

F Ecotone Wildlife, PO Box 76, Inverloch, Vic. 3096, Australia.

G Limestone Coast Landscape Board, 61 Anzac Terrace, Keith, SA 5267, Australia.

H Present address: Faculty of Veterinary and Agricultural Sciences, University of Melbourne, Parkville, Vic. 3052, Australia.

* Correspondence to: dave.forsyth@dpi.nsw.gov.au

Handling Editor: Graham Nugent

Wildlife Research 50(9) 617-631 https://doi.org/10.1071/WR21156
Submitted: 29 October 2021  Accepted: 14 April 2022   Published: 6 July 2022

© 2023 The Author(s) (or their employer(s)). Published by CSIRO Publishing. This is an open access article distributed under the Creative Commons Attribution-NonCommercial-NoDerivatives 4.0 International License (CC BY-NC-ND)

Abstract

Context: Helicopter-based shooting has been widely used to harvest deer or control overabundant populations in Australasia, but the effectiveness and cost of this method as a deer control tool has seldom been evaluated.

Aims: We evaluated the effectiveness and costs of helicopter-based shooting of fallow deer (Dama dama) and chital deer (Axis axis) in eastern Australia by quantifying (1) reductions in density, (2) the relationship between numbers killed per hour and deer density (i.e. the functional response), (3) the costs of control and (4) the effort–outcome and cost–outcome relationships.

Methods: We evaluated the costs and effectiveness of 12 aerial shooting operations aiming to reduce fallow deer (n = 8) or chital deer (n = 4) population densities at nine sites in eastern Australia. Sites were characterised by fragmented woodland, and all but one operation aimed to reduce grazing competition with livestock. We used pre-control population density estimates and operational monitoring data to estimate the costs and outcomes of each operation. We combined data from all operations to estimate the relationship between shooting effort and population reduction, as well as costs associated with different levels of effort.

Key results: Population reductions for operations ranged from 5% to 75% for fallow deer, and from 48% to 88% for chital deer. The greatest population reductions occurred when effort per unit area was greatest, and the largest reductions in deer density occurred when shooting was conducted in consecutive years. The functional response of hourly kills to deer density was best described by a modified Ivlev model, with the asymptotic kill rate estimated to be 50 deer per hour. There was no support for the existence of a prey refuge, that is, a threshold population density below which no deer could be shot. Helicopter charter was the primary cost of helicopter-based shooting programs, followed by labour; firearm and ammunition costs were relatively minor.

Conclusions: Helicopter-based shooting can rapidly reduce deer populations over large geographic areas, but the magnitude of the reduction depends on the effort (hours of shooting) per deer per km2.

Implications: Aerial shooting operations should include a pre-control population survey so that (1) measurable objectives can be established, (2) the likely level of effort and cost required for objectives to be met can be estimated and planned for, and (3) the realised population reduction can be estimated.

Keywords: abundance, aerial gunning, aerial shooting, aerial survey, Bayesian statistics, chital deer, culling, density, effort–outcome, fallow deer, functional response, mark–recapture distance sampling.


References

Ahrestani, FS, Van Langevelde, F, Heitkönig, IMA, and Prins, HHT (2012). Contrasting timing of parturition of chital Axis axis and gaur Bos gaurus in tropical South India – the role of body mass and seasonal forage quality. Oikos 121, 1300–1310.
Contrasting timing of parturition of chital Axis axis and gaur Bos gaurus in tropical South India – the role of body mass and seasonal forage quality.Crossref | GoogleScholarGoogle Scholar |

Amos, M, Baxter, G, Finch, N, Lisle, A, and Murray, P (2014). I just want to count them! Considerations when choosing a deer population monitoring method. Wildlife Biology 20, 362–370.
I just want to count them! Considerations when choosing a deer population monitoring method.Crossref | GoogleScholarGoogle Scholar |

(2021). A situation analysis of commercial wild deer harvesting in Australia with reference to New Zealand. Australian Deer 45, 44–47.

Bengsen AJ, Forsyth DM, Pople AR, Brennan M, Amos M, Leeson M, Cox TE, Gray B, Orgill O, Hampton JO, Haebich K (2022a) ‘AndrewBengsen/Helicopter-based-shooting-of-deer_revised.’ (Zenodo) https://doi.org/10.5281/zenodo.6070097

Bengsen, AJ, Forsyth, DM, Ramsey, DSL, Amos, M, Brennan, M, Pople, AR, Comte, S, and Crittle, T (2022b). Estimating deer density and abundance using spatial mark–resight models with camera trap data. Journal of Mammalogy , gyac016.
Estimating deer density and abundance using spatial mark–resight models with camera trap data.Crossref | GoogleScholarGoogle Scholar |

Bennett M (1979) ‘The venison hunters.’ (A.H. & A.W. Reed: Wellington, New Zealand)

Bentley A (1995) Fallow deer. In ‘The mammals of Australia’. 2nd edn. (Ed. R Strahan) pp. 732–733. (Australian Museum/Reed New Holland: Sydney, NSW, Australia)

Brazeal, JL, Weist, T, and Sacks, BN (2017). Noninvasive genetic spatial capture-recapture for estimating deer population abundance: estimating abundance and density of mule deer. The Journal of Wildlife Management 81, 629–640.
Noninvasive genetic spatial capture-recapture for estimating deer population abundance: estimating abundance and density of mule deer.Crossref | GoogleScholarGoogle Scholar |

Buckland ST, Anderson DR, Burnham KP, Laake JL, Borchers DL, Thomas L (2001) ‘Introduction to distance sampling.’ (Oxford University Press: London, UK)

Buckland ST, Anderson DR, Burnham KP, Laake JL, Borchers DL, Thomas L (2004) ‘Advanced distance sampling: estimating abundance of biological populations.’ (Oxford University Press: London, UK)

Burnham KP, Anderson DR (2002) ‘Model selection and multi-model inference.’ (Springer: New York City, NY, USA)

Burt, ML, Borchers, DL, Jenkins, KJ, and Marques, TA (2014). Using mark–recapture distance sampling methods on line transect surveys. Methods in Ecology and Evolution 5, 1180–1191.
Using mark–recapture distance sampling methods on line transect surveys.Crossref | GoogleScholarGoogle Scholar |

Caravaggi, A, Gatta, M, Vallely, M-C, Hogg, K, Freeman, M, Fadaei, E, Dick, JTA, Montgomery, WI, Reid, N, and Tosh, DG (2018). Seasonal and predator–prey effects on circadian activity of free-ranging mammals revealed by camera traps. PeerJ 6, e5827.
Seasonal and predator–prey effects on circadian activity of free-ranging mammals revealed by camera traps.Crossref | GoogleScholarGoogle Scholar | 30498626PubMed |

Carpio, AJ, Apollonio, M, and Acevedo, P (2021). Wild ungulate overabundance in Europe: contexts, causes, monitoring and management recommendations. Mammal Review 51, 95–108.
Wild ungulate overabundance in Europe: contexts, causes, monitoring and management recommendations.Crossref | GoogleScholarGoogle Scholar |

Caughley G (1980) ‘Analysis of vertebrate populations.’ (Wiley: New York, NY, USA)

Challies, CN (1977). Effects of commercial hunting on red deer densities in the Arawata Valley, South Westland, 1972–76. New Zealand Journal of Forestry Science 7, 263–273.

Challies CN (1985a) Commercial hunting of wild red deer in New Zealand. In ‘Game harvest management’. (Eds SL Beasom, SF Robertson) pp. 279–287. (Caesar Kleberg Wildlife Research Institute: Kingsville, TX, USA)

Challies CN (1985b) Establishment, control, and commercial exploitation of wild deer in New Zealand. In ‘Biology of deer production. Vol. 22’. (Eds PF Fenessey, KR Drew) pp. 23–36. (The Royal Society of New Zealand Bulletin)

Choquenot, D, Hone, J, and Saunders, G (1999). Using aspects of predator-prey theory to evaluate helicopter shooting for feral pig control. Wildlife Research 26, 251–261.
Using aspects of predator-prey theory to evaluate helicopter shooting for feral pig control.Crossref | GoogleScholarGoogle Scholar |

Côté, SD, Rooney, TP, Tremblay, J-P, Dussault, C, and Waller, DM (2004). Ecological impacts of deer overabundance. Annual Review of Ecology, Evolution, and Systematics 35, 113–147.
Ecological impacts of deer overabundance.Crossref | GoogleScholarGoogle Scholar |

Crittle T, Millynn B (2020) Pest animal mapping 2020 final report. NSW Department of Primary Industries, Orange, NSW, Australia.

Cunningham, CX, Perry, GLW, Bowman, DMJS, Forsyth, DM, Driessen, MM, Appleby, M, Brook, BW, Hocking, G, Buettel, JC, French, BJ, Hamer, R, Bryant, SL, Taylor, M, Gardiner, R, Proft, K, Scoleri, VP, Chiu-Werner, A, Travers, T, Thompson, L, Guy, T, and Johnson, CN (2022). Dynamics and predicted distribution of an irrupting ‘sleeper’ population: fallow deer in Tasmania. Biological Invasions 24, 1131–1147.
Dynamics and predicted distribution of an irrupting ‘sleeper’ population: fallow deer in Tasmania.Crossref | GoogleScholarGoogle Scholar |

Davis, NE, Bennett, A, Forsyth, DM, Bowman, DMJS, Lefroy, EC, Wood, SW, Woolnough, AP, West, P, Hampton, JO, and Johnson, CN (2016). A systematic review of the impacts and management of introduced deer (family Cervidae) in Australia. Wildlife Research 43, 515–532.
A systematic review of the impacts and management of introduced deer (family Cervidae) in Australia.Crossref | GoogleScholarGoogle Scholar |

Denwood, MJ (2016). runjags: an R package providing interface utilities, model templates, parallel computing methods and additional distributions for MCMC models in JAGS. Journal of Statistical Software 71, 1–25.
runjags: an R package providing interface utilities, model templates, parallel computing methods and additional distributions for MCMC models in JAGS.Crossref | GoogleScholarGoogle Scholar |

Department for Environment and Water (SA) (2019) ‘Standard operating procedure – aerial shooting of feral animals.’ (Department for Environment and Water (SA): Adelaide, SA, Australia)

Duncan, RP, Forsyth, DM, and Hone, J (2007). Testing the metabolic theory of ecology: allometric scaling exponents in mammals. Ecology 88, 324–333.
Testing the metabolic theory of ecology: allometric scaling exponents in mammals.Crossref | GoogleScholarGoogle Scholar | 17479751PubMed |

Edwards, GP, Saalfeld, K, and Clifford, B (2004). Population trend of feral camels in the Northern Territory, Australia. Wildlife Research 31, 509–517.
Population trend of feral camels in the Northern Territory, Australia.Crossref | GoogleScholarGoogle Scholar |

Ensing, EP, Ciuti, S, de Wijs, FALM, Lentferink, DH, ten Hoedt, A, Boyce, MS, and Hut, RA (2014). GPS based daily activity patterns in European Red Deer and North American Elk (Cervus elaphus): indication for a weak circadian clock in ungulates. PLoS ONE 9, e106997.
GPS based daily activity patterns in European Red Deer and North American Elk (Cervus elaphus): indication for a weak circadian clock in ungulates.Crossref | GoogleScholarGoogle Scholar | 25208246PubMed |

Feral Animal Aerial Shooting Team (2020) ‘The FAAST manual. Version 2.3 (August 2020).’ (National Parks & Wildlife Service, NSW Local Land Services and NSW Department of Primary Industries, on behalf of the NSW Government: Sydney, NSW, Australia)

Forsyth, DM, Wilmshurst, JM, Allen, RB, and Coomes, DA (2010a). Impacts of introduced deer and extinct moa on New Zealand ecosystems. New Zealand Journal of Ecology 34, 48–65.

Forsyth, DM, Allen, RB, Marburg, AE, MacKenzie, DI, and Douglas, MJW (2010b). Population dynamics and resource use of red deer following release from harvesting in New Zealand. New Zealand Journal of Ecology 34, 277–287.

Forsyth, DM, Thomson, C, Hartley, L, MacKenzie, DI, Price, R, Wright, EF, Mortimer, J, Nugent, G, Wilson, L, and Livingstone, P (2011). Long-term changes in the relative abundances of introduced deer in New Zealand estimated from faecal pellet frequencies. New Zealand Journal of Zoology 38, 237–249.
Long-term changes in the relative abundances of introduced deer in New Zealand estimated from faecal pellet frequencies.Crossref | GoogleScholarGoogle Scholar |

Forsyth, DM, Ramsey, DSL, Veltman, CJ, Allen, RB, Allen, WJ, Barker, RJ, Jacobson, CL, Nicol, SJ, Richardson, SJ, and Todd, CR (2013). When deer must die: large uncertainty surrounds changes in deer abundance achieved by helicopter- and ground-based hunting in New Zealand forests. Wildlife Research 40, 447–458.
When deer must die: large uncertainty surrounds changes in deer abundance achieved by helicopter- and ground-based hunting in New Zealand forests.Crossref | GoogleScholarGoogle Scholar |

Forsyth DM, Pople T, Page B, Moriarty A, Ramsey D, Parkes J, Wiebkin A, Lane C (Eds) (2017) ‘2016 National wild deer management workshop proceedings’, Adelaide, SA, Australia, 17–18 November 2016. (Invasive Animals Cooperative Research Centre: Canberra, ACT, Australia) Available at https://pestsmart.org.au/wp-content/uploads/sites/3/2020/06/Proceedings-2016-National-Workshop-on-Deer-Management_FINAL.pdf

Forsyth, DM, Pople, A, Woodford, L, Brennan, M, Amos, M, Moloney, PD, Fanson, B, and Story, G (2019). Landscape-scale effects of homesteads, water, and dingoes on invading chital deer in Australia’s dry tropics. Journal of Mammalogy 100, 1954–1965.
Landscape-scale effects of homesteads, water, and dingoes on invading chital deer in Australia’s dry tropics.Crossref | GoogleScholarGoogle Scholar |

Forsyth, DM, Comte, S, Davis, NE, Bengsen, AJ, Côté, SD, Hewitt, DG, Morellet, N, and Mysterud, A (2022). Methodology matters when estimating deer abundance: a global systematic review and recommendations for improvements. Journal of Wildlife Management 2022, e22207.
Methodology matters when estimating deer abundance: a global systematic review and recommendations for improvements.Crossref | GoogleScholarGoogle Scholar |

Gentle, M, and Pople, A (2013). Effectiveness of commercial harvesting in controlling feral-pig populations. Wildlife Research 40, 459–469.
Effectiveness of commercial harvesting in controlling feral-pig populations.Crossref | GoogleScholarGoogle Scholar |

Graf, W, and Nichols, L (1966). The axis deer in Hawaii. Journal of the Bombay Natural History Society 63, 630–734.

Hampton, JO, Bengsen, AJ, Pople, A, Brennan, M, Leeson, M, and Forsyth, DM (2022). Animal welfare outcomes of helicopter-based shooting of deer in Australia. Wildlife Research 49, 264–273.
Animal welfare outcomes of helicopter-based shooting of deer in Australia.Crossref | GoogleScholarGoogle Scholar |

Hone, J (1990). Predator prey theory and feral pig control, with emphasis on evaluation of shooting from a helicopter. Australian Wildlife Research 17, 123–130.
Predator prey theory and feral pig control, with emphasis on evaluation of shooting from a helicopter.Crossref | GoogleScholarGoogle Scholar |

Hone J (1994) ‘Analysis of vertebrate pest control.’ (Cambridge University Press: Cambridge, UK)

Hone, J (1999). On rate of increase (r): patterns of variation in Australian mammals and the implications for wildlife management. Journal of Applied Ecology 36, 709–718.
On rate of increase (r): patterns of variation in Australian mammals and the implications for wildlife management.Crossref | GoogleScholarGoogle Scholar |

Hone, J, Duncan, RP, and Forsyth, DM (2010). Estimates of maximum annual population growth rates (rm) of mammals and their application in wildlife management. Journal of Applied Ecology 47, 507–514.
Estimates of maximum annual population growth rates (rm) of mammals and their application in wildlife management.Crossref | GoogleScholarGoogle Scholar |

Hone, J, Drake, VA, and Krebs, CJ (2017). The effort–outcomes relationship in applied ecology: evaluation and implications. BioScience 67, 845–852.
The effort–outcomes relationship in applied ecology: evaluation and implications.Crossref | GoogleScholarGoogle Scholar |

Iacona, GD, Sutherland, WJ, Mappin, B, Adams, VM, Armsworth, PR, Coleshaw, T, Cook, C, Craigie, I, Dicks, LV, Fitzsimons, JA, McGowan, J, Plumptre, AJ, Polak, T, Pullin, AS, Ringma, J, Rushworth, I, Santangeli, A, Strwart, A, Tulloch, A, Walsh, JC, and Possingham, HP (2018). Standardized reporting of the costs of management interventions for biodiversity conservation. Conservation Biology 32, 979–988.
Standardized reporting of the costs of management interventions for biodiversity conservation.Crossref | GoogleScholarGoogle Scholar | 30039609PubMed |

Johnson, KA, and Goody, RS (2011). The original Michaelis constant: translation of the 1913 Michaelis–Menten paper. Biochemistry 50, 8264–8269.
The original Michaelis constant: translation of the 1913 Michaelis–Menten paper.Crossref | GoogleScholarGoogle Scholar | 21888353PubMed |

King, AD, Pitman, AJ, Henley, BJ, Ukkola, AM, and Brown, JR (2020). The role of climate variability in Australian drought. Nature Climate Change 10, 177–179.
The role of climate variability in Australian drought.Crossref | GoogleScholarGoogle Scholar |

Laake J, Borchers D, Thomas L, Miller D, Bishop J (2020) mrds: mark–recapture distance sampling. R package version 2.4.4. Available at https://CRAN.R-project.org/package=mrds [Verified 26 August 2021]

Latham, ADM, Cecilia Latham, M, Herries, D, Barron, M, Cruz, J, and Anderson, DP (2018). Assessing the efficacy of aerial culling of introduced wild deer in New Zealand with analytical decomposition of predation risk. Biological Invasions 20, 251–266.
Assessing the efficacy of aerial culling of introduced wild deer in New Zealand with analytical decomposition of predation risk.Crossref | GoogleScholarGoogle Scholar |

Lethbridge MR (2019) Feral deer aerial survey of Gum Lagoon Conservation Park and Surrounds, 2019. Report to Upper South East District. Parks and Regions, Department of Environment, Water and Natural Resources, Adelaide, SA, Australia.

Marsh, H, and Sinclair, DF (1989). Correcting for visibility bias in strip transect aerial surveys of aquatic fauna. The Journal of Wildlife Management 53, 1017–1024.
Correcting for visibility bias in strip transect aerial surveys of aquatic fauna.Crossref | GoogleScholarGoogle Scholar |

McLeod SR (2018) ‘Deer aerial survey logger. v1.0.’ (NSW Department of Primary Industries, Vertebrate Pest Research Unit: Orange, NSW, Australia)

McShea WJ, Underwood HB, Rappole JH (Eds) (1997) ‘The science of overabundance: deer ecology and population management.’ (Smithsonian Institution Press: Washington, DC, USA)

Miller, DL, Burt, ML, Rexstad, EA, and Thomas, L (2013). Spatial models for distance sampling data: recent developments and future directions. Methods in Ecology and Evolution 4, 1001–1010.
Spatial models for distance sampling data: recent developments and future directions.Crossref | GoogleScholarGoogle Scholar |

Moloney, PD, Forsyth, DM, Ramsey, DSL, Perry, M, McKay, M, Gormley, AM, Kappers, B, and Wright, EF (2021). Occupancy and relative abundances of introduced ungulates on New Zealand’s public conservation land 2012–2018. New Zealand Journal of Ecology 45, 3437.
Occupancy and relative abundances of introduced ungulates on New Zealand’s public conservation land 2012–2018.Crossref | GoogleScholarGoogle Scholar |

Moriarty, A (2004). The liberation, distribution, abundance and management of wild deer in Australia. Wildlife Research 31, 291–299.
The liberation, distribution, abundance and management of wild deer in Australia.Crossref | GoogleScholarGoogle Scholar |

Morriss, GA, Parkes, JP, and Nugent, G (2020). Effects of aerial 1080 operations on deer populations in New Zealand. New Zealand Journal of Ecology 44, 3417.
Effects of aerial 1080 operations on deer populations in New Zealand.Crossref | GoogleScholarGoogle Scholar |

Naidoo, R, Balmford, A, Ferraro, PJ, Polasky, S, Ricketts, TH, and Rouget, M (2006). Integrating economic costs into conservation planning. Trends in Ecology & Evolution 21, 681–687.
Integrating economic costs into conservation planning.Crossref | GoogleScholarGoogle Scholar |

Nugent G, Asher GW (2021) Family Cervidae: Dama dama. In ‘The handbook of New Zealand mammals’. 3rd edn. (Eds CM King, DM Forsyth) pp. 447–527. (CSIRO: Melbourne, Vic., Australia)

Nugent, G, and Choquenot, D (2004). Comparing cost-effectiveness of commercial harvesting, state-funded culling, and recreational deer hunting in New Zealand. Wildlife Society Bulletin 32, 481–492.
Comparing cost-effectiveness of commercial harvesting, state-funded culling, and recreational deer hunting in New Zealand.Crossref | GoogleScholarGoogle Scholar |

Nugent G, Forsyth DM (2021) Family Cervidae: Cervus elaphus. In ‘The handbook of New Zealand mammals’. 3rd edn. (Eds CM King, DM Forsyth) pp. 447–527. (CSIRO: Melbourne, Vic., Australia)

Nugent, G, and Yockney, I (2004). Fallow deer deaths during aerial-1080 poisoning of possums in the Blue Mountains, Otago, New Zealand. New Zealand Journal of Zoology 31, 185–192.
Fallow deer deaths during aerial-1080 poisoning of possums in the Blue Mountains, Otago, New Zealand.Crossref | GoogleScholarGoogle Scholar |

Nugent, G, Parkes, JP, and Tustin, KG (1987). Changes in the density and distribution of red deer and wapiti in northern Fiordland. New Zealand Journal of Ecology 10, 11–21.

Parkes, JP, Nugent, G, and Warburton, B (1996). Commercial exploitation as a pest control tool for introduced mammals in New Zealand. Wildlife Biology 2, 171–177.
Commercial exploitation as a pest control tool for introduced mammals in New Zealand.Crossref | GoogleScholarGoogle Scholar |

Peterson, MK, Foley, AM, Tri, AN, Hewitt, DG, DeYoung, RW, DeYoung, CA, and Campbell, TA (2020). Mark-recapture distance sampling for aerial surveys of ungulates on rangelands. Wildlife Society Bulletin 44, 713–723.
Mark-recapture distance sampling for aerial surveys of ungulates on rangelands.Crossref | GoogleScholarGoogle Scholar |

Plummer M (2003) JAGS: a program for analysis of Bayesian graphical models using Gibbs sampling. In ‘Proceedings of the 3rd international workshop on distributed statistical computing, Vol. 124, no. 125’. (Eds K Hornik, F Leisch, A Zelleis) pp. 1–10. (Technische Universität Wien: Vienna, Austria)

Ramsey DSL, Pacioni C, Hill E (2019) Abundance and population genetics of Hog Deer (Axis porcinus) in Victoria. Technical Report Series No. 303. Arthur Rylah Institute for Environmental Research, Department of Environment, Land, Water and Planning, Melbourne, Vic., Australia.

R Core Team (2020) ‘R: a language and environment for statistical computing.’ Version 4.0.3. (R Foundation for Statistical Computing) Available at https://www.R-project.org/ [Verified 26 August 2021]

Rao, TR (2000). A curve for all reasons. Resonance 5, 85–90.
A curve for all reasons.Crossref | GoogleScholarGoogle Scholar |

Senate Select Committee on Animal Welfare (1991) Culling of large feral animals in the northern territory. Report by the Senate Select Committee on Animal Welfare. Senate Publications Unit, Parliament House, Canberra, ACT, Australia.

Sharp T (2012) ‘Standard operating procedure GOA002: aerial shooting of feral goats.’ (Centre for Invasive Species Solutions: Canberra, ACT, Australia)

Standing Committee on Agriculture, Animal Health Committee (2002) Model code of practice for the welfare of animals. Feral Livestock Animals. Destruction or Capture, Handling and Marketing. SCARM Report 34. CSIRO, Melbourne, Vic., Australia.

Strindberg, S, and Buckland, ST (2004). Zigzag survey designs in line transect sampling. Journal of Agricultural, Biological, and Environmental Statistics 9, 443.
Zigzag survey designs in line transect sampling.Crossref | GoogleScholarGoogle Scholar |

Thomas, L, Buckland, ST, Rexstad, EA, Laake, JL, Strindberg, S, Hedley, SL, Bishop, JRB, Marques, TA, and Burnham, KP (2010). Distance software: design and analysis of distance sampling surveys for estimating population size. Journal of Applied Ecology 47, 5–14.
Distance software: design and analysis of distance sampling surveys for estimating population size.Crossref | GoogleScholarGoogle Scholar | 20383262PubMed |

Tracey J, Fleming PJS (2016) ‘Techniques for aerial surveys of wildlife: reference manual.’ 4th edn. (Vertebrate Pest Research Unit, NSW Department of Primary Industries: Orange, NSW, Australia)

Tracey J, Fleming PJS, McLeod S, Purcell B (2016) ‘Techniques for aerial surveys of wildlife: training manual.’ 4th edn. (Vertebrate Pest Research Unit, NSW Department of Primary Industries: Orange, NSW, Australia)

Tustin, KG, and Challies, CN (1978). The effects of hunting on the numbers and group sizes of Himalayan thar (Hemitragus jemlahicus) in Carneys Creek, Rangitata catchment. New Zealand Journal of Ecology 1, 153–157.

van Dyck S, Strahan R (2008) ‘The mammals of Australia.’ (New Holland Publishers: Sydney, NSW, Australia)

Veltman, CJ, and Pinder, DN (2001). Brushtail possum mortality and ambient temperatures following aerial poisoning using 1080. The Journal of Wildlife Management 65, 476–481.
Brushtail possum mortality and ambient temperatures following aerial poisoning using 1080.Crossref | GoogleScholarGoogle Scholar |

Warburton B, Anderson DP, Nugent G (2018) Economic aspects of New Zealand’s wild venison recovery industry. In ‘Advances in conservation through sustainable use of wildlife. Proceedings of a conference held in Brisbane, Australia, 20 August − 1 September 2016’. (Eds G Baxter, N Finch, P Murray) pp. 265–271. (University of Queensland: Gatton, Qld, Australia)

Watter, K, Baxter, GS, Pople, A, and Murray, PJ (2020). Dietary overlap between cattle and chital in the Queensland dry tropics. The Rangeland Journal 42, 221–225.
Dietary overlap between cattle and chital in the Queensland dry tropics.Crossref | GoogleScholarGoogle Scholar |