
TY  - JOUR
A1  - Smith, M 
T1  - Behaviour of the Koala, Phascolarctos cinereus (Goldfuss), in Captivity IV.* Scent-Marking
JF  - Wildlife Research
JO  - Wildl. Res.
Y1  - 1980
VL  - 7
IS  - 1
SP  - 35
EP  - 40
KW  - 
UR  - https://doi.org/10.1071/WR9800035 
N2  - Male koalas scent-mark by grasping a vertical object, generally a tree, and rubbing the sternal gland against it. The behaviour first appears at 3 y of age, and reaches a peak a year or two later.  It is frequently performed in conjunction with bellowing, especially in response to agonistic encounters or the bellow of a rival male, and often by itself in circumstances where a bellow might be expected.  It is sometimes a response to unfamiliar surroundings or objects, and otherwise is performed by a male moving about a pen. Scent-marking may primarily be a response to unfamiliarity which, having become associated with displaced aggression, has been entrained to the bellowing drive. 
ER  -

