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^A US Geological Survey, Great Lakes Science Center, Lake Erie Biological Station, 6100 Columbus Avenue, Sandusky, OH 44870, USA.

^B Savannah River Ecology Laboratory, Warnell School of Forestry and Natural Resources, University of Georgia, PO Drawer E, Aiken, SC 29802, USA.

^C Center for Systems Integration and Sustainability, Department of Fisheries and Wildlife, Michigan State University, 115 Manly Miles Building, 1405 S. Harrison Road, East Lansing, MI 48823, USA.

^D Sandusky Fisheries Research Station, Division of Wildlife, Ohio Department of Natural Resources, 305 East Shoreline Drive, Sandusky, OH 44870, USA.

^E Corresponding author. Email: rkraus@usgs.gov