Management of damage by elk (Cervus elaphus) in North America: a review
W. David Walter A , Michael J. Lavelle A , Justin W. Fischer A , Therese L. Johnson B , Scott E. Hygnstrom C and Kurt C. VerCauteren A D
+ Author Affiliations
- Author Affiliations
A United States Department of Agriculture, Animal and Plant Health Inspection Services, National Wildlife Research Center, 4101 LaPorte Ave., Fort Collins, CO 80521, USA.
B Rocky Mountain National Park, Estes Park, CO 80517, USA.
C University of Nebraska, School of Natural Resources, 415 Hardin Hall, 3310 Holdrege Street, Lincoln, NE 68583, USA.
D Corresponding author. Email: Kurt.C.VerCauteren@aphis.usda.gov
Wildlife Research 37(8) 630-646 https://doi.org/10.1071/WR10021
Submitted: 11 February 2010 Accepted: 16 September 2010 Published: 22 December 2010
Abstract
Abundant populations of elk (Cervus elaphus) are cherished game in many regions of the world and also cause considerable human–wildlife conflicts through depredation on agriculture and speciality crops, lack of regeneration to native ecosystems, collisions with vehicles and transmission of disease between free-ranging and farmed hoofstock. Management of elk varies, depending on current and historical agency objectives, configuration of the landscapes elk occupy, public perception, population density and behaviour of elk. Selection of the method to manage elk often requires knowledge of timing of impacts, duration relief from elk damage is desired, cost-effectiveness of management activities, tolerance of impacts, public perception of management strategies and motivation or habituation of elk to determine the likelihood of success for a proposed management action. We reviewed methods that are available to control abundant populations of elk that include lethal (e.g. hunting, sharpshooting) and non-lethal (e.g. fertility control, frightening) options. We promote an integrated approach that incorporates the timely use of a variety of cost-effective methods to reduce impacts to tolerable levels. Lethal options that include regulated hunting, sharpshooting and aerial gunning vary by likelihood of success, duration needed for population reduction, cost to implement reduction and public perceptions. Several non-lethal options are available to affect population dynamics directly (e.g. fertility control, translocation), protect resources from damage (e.g. fences, repellents) or influence space use of elk on a regular basis (e.g. harassment, frightening, herding dogs, humans). Public perception should be considered by agencies that are looking for feasible methods to control populations of elk. Disturbance to residents or visitors of public property may influence methods of management employed. Future research should explore the duration of harassment needed to avert elk from sensitive areas and costs to implement such programs. Several methods in our review were implemented on deer and additional research on elk and other cervids in conflict with human interests would provide a much needed component to our understanding of management methods available for ungulate species.
Additional keywords.: Cervus elaphus, damage, ecosystem, elk, fence, fertility control, frightening, habituation, hazing, lethal, non-lethal, regeneration.
References
American Veterinary Medical Association (AVMA) (2007). ‘AVMA Guidelines on Euthanasia.’ Available at http://www.avma.org/issues/animal_welfare/euthanasia.pdf [accessed 15 June 2010].Andelt, W. F., Burnham, K. P., and Manning, J. A. (1991). Relative effectiveness of repellents for reducing mule deer damage. The Journal of Wildlife Management 55, 341–347.
| Relative effectiveness of repellents for reducing mule deer damage.Crossref | GoogleScholarGoogle Scholar |
Andelt, W. F., Baker, D. L., and Burnham, K. P. (1992). Relative preference of captive cow elk for repellent-treated diets. The Journal of Wildlife Management 56, 164–173.
| Relative preference of captive cow elk for repellent-treated diets.Crossref | GoogleScholarGoogle Scholar |
Andelt, W. F., Burnham, K. P., and Baker, D. L. (1994). Effectiveness of capsaicin and bitrex repellents for deterring browsing by mule deer. The Journal of Wildlife Management 58, 330–335.
| Effectiveness of capsaicin and bitrex repellents for deterring browsing by mule deer.Crossref | GoogleScholarGoogle Scholar |
Angradi, T. R., and Tzilkowski, W. M. (1987). Preliminary testing of a selenium-based systemic deer browse repellent. Proceedings of the Eastern Wildlife Damage Control Conference 3, 102–107.
Ayotte, J. B., Parker, K. L., Arocena, J. M., and Gillingham, M. P. (2006). Chemical composition of lick soils: functions of soil ingestion by four ungulate species. Journal of Mammalogy 87, 878–888.
| Chemical composition of lick soils: functions of soil ingestion by four ungulate species.Crossref | GoogleScholarGoogle Scholar |
Baker, D. L., and Hobbs, N. T. (1982). Composition and quality of elk summer diets in Colorado. The Journal of Wildlife Management 46, 694–703.
| Composition and quality of elk summer diets in Colorado.Crossref | GoogleScholarGoogle Scholar |
Baker, D. L., Andelt, W. F., Burnham, K. P., and Shepperd, W. D. (1999). Effectiveness of Hot Sauce® and Deer Away® repellents for deterring elk browsing of aspen sprouts. The Journal of Wildlife Management 63, 1327–1336.
| Effectiveness of Hot Sauce® and Deer Away® repellents for deterring elk browsing of aspen sprouts.Crossref | GoogleScholarGoogle Scholar |
Baker, D. L., Wild, M. A., Hussain, M. D., Dunn, R. L., and Nett, T. M. (2005). Evaluation of remotely delivered leuprolide acetate as a contraceptive agent in female elk (Cervus elaphus nelsoni). Journal of Wildlife Diseases 41, 758–767.
| 1:CAS:528:DC%2BD28XitFOqurc%3D&md5=fb0128478f9fd4aaf687eeb386e943fcCAS | 16456165PubMed |
Bashore, T. L., Tzilkowski, W. M., and Bellis, E. D. (1985). Analysis of deer–vehicle collision sites in Pennsylvania. The Journal of Wildlife Management 49, 769–774.
| Analysis of deer–vehicle collision sites in Pennsylvania.Crossref | GoogleScholarGoogle Scholar |
Bauman, P. J., Jenks, J. A., and Roddy, D. E. (1999). Evaluating techniques to monitor elk movement across fence lines. Wildlife Society Bulletin 27, 344–352.
Bayoumi, M. A., and Smith, A. D. (1976). Response of big game winter range vegetation to fertilization. Journal of Range Management 29, 44–48.
| Response of big game winter range vegetation to fertilization.Crossref | GoogleScholarGoogle Scholar |
Beck, J. L., and Peek, J. M. (2005). Diet composition, forage selection and potential for forage competition among elk, deer and livestock on aspen–sagebrush summer range. Rangeland Ecology and Management 58, 135–147.
| Diet composition, forage selection and potential for forage competition among elk, deer and livestock on aspen–sagebrush summer range.Crossref | GoogleScholarGoogle Scholar |
Belant, J. L., Seamans, T. W., and Dwyer, C. P. (1996). Evaluation of propane exploders as white-tailed deer deterrents. Crop Protection 15, 575–578.
| Evaluation of propane exploders as white-tailed deer deterrents.Crossref | GoogleScholarGoogle Scholar |
Belant, J. L., Seamans, T. W., and Tyson, L. A. (1998). Evaluation of electronic frightening devices as white-tailed deer deterrents. Proceedings of the Vertebrate Pest Conference 18, 107–110.
Belovsky, G. E., and Jordan, P. A. (1981). Sodium dynamics and adaptations of a moose population. Journal of Mammalogy 62, 613–621.
| Sodium dynamics and adaptations of a moose population.Crossref | GoogleScholarGoogle Scholar |
Beringer, J., Hansen, L. P., Heinen, R. A., and Geissman, N. F. (1994). Use of dogs to reduce damage by deer to a white pine plantation. Wildlife Society Bulletin 22, 627–632.
Beringer, J., Hansen, L. P., Wilding, W., Fischer, J., and Sheriff, S. L. (1996). Factors affecting capture myopathy in white-tailed deer. The Journal of Wildlife Management 60, 373–380.
| Factors affecting capture myopathy in white-tailed deer.Crossref | GoogleScholarGoogle Scholar |
Beringer, J., Hansen, L. P., Demand, J. A., Sartwell, J., Wallendorf, M., and Mange, R. (2002). Efficacy of translocation to control urban deer in Missouri: costs, efficiency, and outcome. Wildlife Society Bulletin 30, 767–774.
Beringer, J., VerCauteren, K. C., and Millspaugh, J. J. (2003). Evaluation of an animal activated scarecrow and a monofilament fence for reducing deer use of soybean fields. Wildlife Society Bulletin 31, 492–498.
Beschta, R. L., and Ripple, W. J. (2006). River channel dynamics following extirpation of wolves in northwestern Yellowstone National Park, USA. Earth Surface Processes and Landforms 31, 1525–1539.
| River channel dynamics following extirpation of wolves in northwestern Yellowstone National Park, USA.Crossref | GoogleScholarGoogle Scholar |
Bienen, L., and Tabor, G. (2006). Applying an ecosystem approach to brucellosis control: can an old conflict between wildlife and agriculture be successfully managed? Frontiers in Ecology and the Environment 4, 319–327.
| Applying an ecosystem approach to brucellosis control: can an old conflict between wildlife and agriculture be successfully managed?Crossref | GoogleScholarGoogle Scholar |
Biggs, J., Sherwood, S., Michalak, S., Hansen, L., and Bare, C. (2004). Animal-related vehicle accidents at the Los Alamos Nationa Laboratory, New Mexico. The Southwestern Naturalist 49, 384–394.
| Animal-related vehicle accidents at the Los Alamos Nationa Laboratory, New Mexico.Crossref | GoogleScholarGoogle Scholar |
Bissonette, J. A., and Cramer, P. C. (2008). Evaluation of the use and effectiveness of wildlife crossings. National Cooperative Highway Research Program Report 615. Transportation Research Board, Washington, DC.
Blackwell, B. F., Bernhardt, G. E., and Dolbeer, R. A. (2002). Lasers as nonlethal avian repellents. The Journal of Wildlife Management 66, 250–258.
| Lasers as nonlethal avian repellents.Crossref | GoogleScholarGoogle Scholar |
Bomford, M., and O’Brien, P. H. (1990). Sonic deterrents in animal damage control: a review of device tests and effectiveness. Wildlife Society Bulletin 18, 411–422.
Boyce, M. S. (1989). ‘The Jackson Elk Herd: Intensive Wildlife Management in North America.’ (Cambridge University Press: New York.)
Bradford, J., and Hobbs, N. T. (2006). Analysis of options for elk population management in Rocky Mountain National Park – results of additional scenarios. Report to the National Park Service. Rocky Mountain National Park, Estes Park, CO.
Bradford, J. B., and Hobbs, N. T. (2008). Regulating overabundant ungulate populations: an example for elk in Rocky Mountain National Park, Colorado. Journal of Environmental Management 86, 520–528.
| Regulating overabundant ungulate populations: an example for elk in Rocky Mountain National Park, Colorado.Crossref | GoogleScholarGoogle Scholar | 17276577PubMed |
Brelsford, M. J., Peek, J. M., and Murray, G. A. (1998). Effects of grazing by wapiti on winter wheat in northern Idaho. Wildlife Society Bulletin 26, 203–208.
Brook, R. K., and McLachlan, S. M. (2006). Factors influencing farmers’ concerns regarding bovine tuberculosis in wildlife and livestock around Riding Mountain National Park. Journal of Environmental Management 80, 156–166.
| Factors influencing farmers’ concerns regarding bovine tuberculosis in wildlife and livestock around Riding Mountain National Park.Crossref | GoogleScholarGoogle Scholar | 16359775PubMed |
Brookshire, J., Kauffman, J. B., Lytjen, D., and Otting, N. (2002). Cumulative effects of wild ungulate and livestock herbivory on riparian willows. Oecologia 132, 559–566.
| Cumulative effects of wild ungulate and livestock herbivory on riparian willows.Crossref | GoogleScholarGoogle Scholar |
Bryant, B. K., and Ishmael, W. E. (1991). Movement and mortality patterns of resident and translocated suburban white-tailed deer. In ‘Proceedings of the National Institute for Urban Wildlife Symposium’. (Eds L. W. Adams and D. L. Leedy.) pp. 53–58. (National Institute for Urban Wildlife: Columbia, MD.)
Bryant, L. D., Thomas, J. W., and Rowland, M. M. (1993). Techniques to construct New Zealand elk-proof fence. United States Department of Agriculture, General Technical Report PNW-GTR-313. United States Forest Service, Pacific Northwest Research Station, Portland, OR.
Bunnell, S. D., Wolfe, M. L., Brunson, M. W., and Potter, D. R. (2002). Recreational use of elk. In ‘North American Elk: Ecology and Management’. (Eds D. E. Toweill and J. W. Thomas.) pp. 701–747. (Smithsonian Institute Press: Washington, DC.)
Burcham, M., Edge, W. D., and Marcum, C. L. (1999). Elk use of private land refuges. Wildlife Society Bulletin 27, 833–839.
Byrne, E. A. (1989). Experimental applications of high-tensile wire and other fencing to control big game damage in northwest Colorado. Proceedings of the Great Plains Wildlife Damage Control Workshop 9, 109–115.
Campbell, D. L., Evans, J., and Hartman, G. B. (1988). Evaluation of seedling protection materials in western Oregon. Technical note OR-5. United States Department of the Interior, Bureau of Land Management, Portland, OR.
Case, G. W. (1938). The use of salt in controlling the distribution of game. The Journal of Wildlife Management 2, 79–81.
| The use of salt in controlling the distribution of game.Crossref | GoogleScholarGoogle Scholar |
Case, R. L., and Kauffman, J. B. (1997). Wild ungulate influences on the recovery of willows, black cottonwood and thin-leaf alder following cessation of cattle grazing in northeastern Oregon. Northwest Science 71, 115–126.
Cassirer, E. F., Freddy, D. J., and Ables, E. D. (1992). Elk reponses to disturbance by cross-country skiers in Yellowstone National Park. Wildlife Society Bulletin 20, 375–381.
Castelli, P. M., and Sleggs, S. E. (2000). Efficacy of border collies to control nuisance Canada geese. Wildlife Society Bulletin 28, 385–392.
Chase, L. C., and Decker, D. J. (1998). Citizen attitude toward elk and participation in elk management: a case study in Evergreen, Colorado. Human Dimensions of Wildlife 3, 55–56.
Clarke, J. A., Olmsted, C. E., and Larkins, K. (1994). Patterns of habitat use, movement, and distribution of elk adjacent to the eastern boundary of Rocky Mountain National Park. Summary report, solicitation #1443RP1520920002. University of Northern Colorado, Greeley, CO.
Clevenger, A. P., Chruszcz, B., and Gunson, K. E. (2001). Highway mitigation fencing reduces wildife–vehicle collisions. Wildlife Society Bulletin 29, 646–653.
Conner, M. M., White, G. C., and Freddy, D. J. (2001). Elk movement in response to early-season hunting in northwest Colorado. The Journal of Wildlife Management 65, 926–940.
| Elk movement in response to early-season hunting in northwest Colorado.Crossref | GoogleScholarGoogle Scholar |
Conner, M. M., Baker, D. L., Wild, M. A., Powers, J. G., Hussain, M. D., Dunn, R. L., and Nett, T. M. (2007). Fertility control in free-ranging elk using gonadotropin-releasing hormone agonist leuprolide: effects on reproduction, behavior and body condidion. The Journal of Wildlife Management 71, 2346–2356.
| Fertility control in free-ranging elk using gonadotropin-releasing hormone agonist leuprolide: effects on reproduction, behavior and body condidion.Crossref | GoogleScholarGoogle Scholar |
Conover, M. R. (1984). Effectiveness of repellents in reducing deer damage in nurseries. Wildlife Society Bulletin 12, 399–404.
Conover, M. (2002). ‘Resolving Human–wildlife Conflicts: The Science of Wildlife Damage Management.’ (Lewis Publishers: Boca Raton, FL.)
Cooney, R. F. (1952). Elk problems in Montana. Journal of Range Management 5, 3–7.
| Elk problems in Montana.Crossref | GoogleScholarGoogle Scholar |
Cooper, A. B., Stewart, F., Unsworth, J. W., Kuck, L., McArthur, T. J., and Rachael, J. S. (2002). Incorporating economic impacts into wildlife management decisions. Wildlife Society Bulletin 30, 565–574.
Coppinger, R., Lorenz, J., and Coppinger, L. (1987). New uses of livestock guarding dogs to reduce agriculture/wildlife conflicts. Proceedings of the Eastern Wildlife Damage Control Conference 3, 253–259.
Coughenour, M. B., and Singer, F. J. (1996). Elk population processes in Yellowstone National Park under the policy of natural regulation. Ecological Applications 6, 573–593.
| Elk population processes in Yellowstone National Park under the policy of natural regulation.Crossref | GoogleScholarGoogle Scholar |
Craven, S. R., and Hygnstrom, S. E. (1994). Deer. In ‘Prevention and Control of Wildlife Damage’. (Eds S. Hygnstrom, R. Timm and G. Larsen.) pp. D-25–D-40. (University of Nebraska Press: Lincoln, NE.)
Creel, S., Winnie, J., Maxwell, B., Hamlin, K., and Creel, M. (2005). Elk alter habitat selection as an antipredator response to wolves. Ecology 86, 3387–3397.
| Elk alter habitat selection as an antipredator response to wolves.Crossref | GoogleScholarGoogle Scholar |
Cromwell, J. A., Warren, R. J., and Henderson, D. W. (1999). Live-capture and small-scale relocation of urban deer on Hilton Head Island, South Carolina. Wildlife Society Bulletin 27, 1025–1031.
Cross, P. C., Edwards, W. H., Scurlock, B. M., Maichak, E. J., and Rogerson, J. D. (2007). Effects of management and climate on elk brucellosis in the greater Yellowstone ecosystem. Ecological Applications 17, 957–964.
| Effects of management and climate on elk brucellosis in the greater Yellowstone ecosystem.Crossref | GoogleScholarGoogle Scholar | 17555209PubMed |
Cross, P. C., Heisey, D. M., Scurlock, B. M., Edwards, W. H., Ebinger, M. R., and Brennan, A. (2010). Mapping brucellosis increases relative to elk density using hierarchical Bayesian models. PLoS ONE 5, e10322.
| Mapping brucellosis increases relative to elk density using hierarchical Bayesian models.Crossref | GoogleScholarGoogle Scholar | 20428240PubMed |
Curtis, P. D., and Rieckenberg, R. (2005). Use of confined dogs for reducing deer damage to apple orchards. Proceedings of the Wildlife Damage Management Conference 11, 149–158.
D’Angelo, G. J., Warren, R. J., Miller, K. V., Gallagher, G. R., and Valitzski, S. A. (2007). Development and evaluation of devices designed to minimize deer–vehicle collisions. Final Project Report, Georgia Department of Transportation. University of Georgia, Athens, GA.
Dalke, P. D., Beeman, R. D., Kindel, F. J., Robel, R. J., and Williams, T. R. (1965). Use of salt by elk in Idaho. The Journal of Wildlife Management 29, 319–332.
| Use of salt by elk in Idaho.Crossref | GoogleScholarGoogle Scholar |
Daniels, M. J., Hutchings, M. R., and Greig, A. (2003). The risk of disease transmission to livestock posed by contamination of farm stored feed by wildlife excreta. Epidemiology and Infection 130, 561–568.
| 1:STN:280:DC%2BD3s3pvVSltg%3D%3D&md5=25e08e30dc36807ce78cf4bd8941fa35CAS | 12825742PubMed |
Dean, R., Gocke, M., Holz, B., Kilpatrick, S., Kreeger, T. J., Scurlock, B., Smith, S., Thorne, E. T., and Werbelow, S. (2004). ‘Elk Feedgrounds in Wyoming.’ (Wyoming Department of Game and Fish: Jackson, WY.)
Deblinger, R. D., Rimmer, D. W., Vaske, J. J., Vecellio, G. M., and Donnelly, M. P. (1993). Ecological benefits and hunter acceptance of a controlled deer hunt in coastal Massachusetts. Northeast Wildlife 50, 11–21.
DeNicola, A. J., and Williams, S. C. (2008). Sharpshooting suburban white-tailed deer reduces deer–vehicle collisions. Human–Wildlife Conflicts 2, 28–33.
DeNicola, A. J., Weber, S. J., Bridges, C. A., and Stokes, J. L. (1997). Nontraditional techniques for management of overabundant deer populations. Wildlife Society Bulletin 25, 496–499.
DeNicola, A. J., Vercauteren, K. C., Curtis, P. D., and Hygnstrom, S. E. (2000). ‘Managing White-tailed Deer in Suburban Environments – A Technical Guide.’ (Cornell Cooperative Extension: Ithaca, NY.)
DeNicola, A. J., Etter, D. R., and Almendinger, T. (2008). Demographics of non-hunted white-tailed deer populations in suburban areas. Human–Wildlife Conflicts 2, 102–109.
Dexter, N. (1996). The effect of an intensive shooting exercise from a helicopter on the behaviour of surviving feral pigs. Wildlife Research 23, 435–441.
| The effect of an intensive shooting exercise from a helicopter on the behaviour of surviving feral pigs.Crossref | GoogleScholarGoogle Scholar |
Didier, K. A., and Porter, W. F. (1999). Large-scale assessment of potential habitat to restore elk to New York State. Wildlife Society Bulletin 27, 409–418.
Dieni, J. S., Smith, R. L., Rogers, R. L., and Anderson, S. H. (2000). Effects of ungulate browsing on aspen regeneration in northwestern Wyoming. Intermountain Journal of Sciences 6, 49–55.
Doenier, P. B., DelGiudice, G. D., and Riggs, M. R. (1997). Effects of winter supplemental feeding on browse consumption of white-tailed deer. Wildlife Society Bulletin 25, 235–243.
Dolbeer, R. A. (1998). Population dynamics: the foundation of wildlife damage management for the 21st century. Proceedings of the Vertebrate Pest Conference 18, 2–11.
Donovan, G., and Champ, P. (2009). The economic benefits of elk viewing at the Jewell Meadows Wildlife Area in Oregon. Human Dimensions of Wildlife 14, 51–60.
| The economic benefits of elk viewing at the Jewell Meadows Wildlife Area in Oregon.Crossref | GoogleScholarGoogle Scholar |
Edge, W. D., and Marcum, C. L. (1985). Movements of elk in relation to logging disturbances. The Journal of Wildlife Management 49, 926–930.
| Movements of elk in relation to logging disturbances.Crossref | GoogleScholarGoogle Scholar |
Etter, R. P., and Drew, M. L. (2006). Brucellosis in elk of eastern Idaho. Journal of Wildlife Diseases 42, 271–278.
| 16870849PubMed |
Fagerstone, K. A., and Clay, W. H. (1997). Overview of USDA animal damage control efforts to manage overabundant deer. Wildlife Society Bulletin 25, 413–417.
Fagerstone, K. A., Coffey, M. A., Curtis, P. D., Dolbeer, R. A., Killian, G. J., Miller, L. A., and Wilmot, L. M. (2002). Wildlife fertility control. The Wildlife Society Technical Review 02-2. The Wildlife Society, Bethesda, MD.
Fargione, M. J., Curtis, P. D., and Richmond, M. E. (1991). Resistance of woody ornamental plants to deer damage. Wildlife damage management program fact sheet. Cornell Cooperative Extension, Ithaca, NY.
Farnsworth, M. L., Wolfe, L. L., Hobbs, N. T., Burnham, K. P., Williams, E. S., Theobald, D. M., Conner, M. M., and Miller, M. W. (2005). Human land use influences chronic wasting disease prevalence in mule deer. Ecological Applications 15, 119–126.
| Human land use influences chronic wasting disease prevalence in mule deer.Crossref | GoogleScholarGoogle Scholar |
Feldhamer, G. A., Gates, J. E., Harman, D. M., Loranger, A. J., and Dixon, K. R. (1986). Effects of interstate highway fencing on white-tailed deer activity. The Journal of Wildlife Management 50, 497–503.
| Effects of interstate highway fencing on white-tailed deer activity.Crossref | GoogleScholarGoogle Scholar |
Frair, J. L., Merrill, E. H., Allen, J. R., and Boyce, M. S. (2007). Know thy enemy: experience affects elk translocation success in risky landscapes. The Journal of Wildlife Management 71, 541–554.
| Know thy enemy: experience affects elk translocation success in risky landscapes.Crossref | GoogleScholarGoogle Scholar |
Fricke, K. A., Cover, M. A., Hygnstrom, S. E., Groepper, S. R., Genoways, H. H., Hams, K. M., and VerCauteren, K. C. (2008). Historic and recent distributions of elk in Nebraska. Great Plains Research 18, 189–204.
Frid, A., and Dill, L. (2002). Human-caused disturbance stimuli as a form of predation risk. Conservation Ecology 6, 1–11.
Gage, E., and Cooper, D. (2009). A study of the effectiveness of a fence design in excluding elk and moose but allowing the movement of other wildlife. Final Research Report, RM-CESU Cooperative Agreement Number H1200040001. Colorado State University, Fort Collins, CO.
Gehring, T. M., Vercauteren, K. C., and Landry, J. (2010). Livestock protection dogs in the 21st century: is an ancient tool relevant to modern conservation challenges? Bioscience 60, 299–308.
| Livestock protection dogs in the 21st century: is an ancient tool relevant to modern conservation challenges?Crossref | GoogleScholarGoogle Scholar |
Gilsdorf, J. M., Hygnstrom, S. E., and VerCauteren, K. C. (2002). Use of frightening devices wildlife damage management. Integrated Pest Management Reviews 7, 29–45.
| Use of frightening devices wildlife damage management.Crossref | GoogleScholarGoogle Scholar |
Gilsdorf, J. M., Hygnstrom, S. E., VerCauteren, K. C., and Blankenship, E. E. (2004a). Propane exploders and electronic guards were ineffective at reducing deer damage in corn fields. Wildlife Society Bulletin 32, 524–531.
| Propane exploders and electronic guards were ineffective at reducing deer damage in corn fields.Crossref | GoogleScholarGoogle Scholar |
Gilsdorf, J. M., Hygnstrom, S. E., VerCauteren, K. C., Clements, G. M., Blankenship, E. E., and Engeman, R. M. (2004b). Evaluation of a deer-activated bio-acoustic frightening device for reducing deer damage in cornfields. Wildlife Society Bulletin 32, 515–523.
| Evaluation of a deer-activated bio-acoustic frightening device for reducing deer damage in cornfields.Crossref | GoogleScholarGoogle Scholar |
Goddard, P. J., Summers, R. W., Macdonald, A. J., Murray, C., and Fawcett, A. R. (2001). Behavioural responses of red deer to fences of five different designs. Applied Animal Behaviour Science 73, 289–298.
| Behavioural responses of red deer to fences of five different designs.Crossref | GoogleScholarGoogle Scholar | 11434963PubMed |
Green, J. S., and Woodruff, R. A. (1990). Livestock guarding dogs: protecting sheep from predators. Agriculture Information Bulletin, Number 588. Extension Service and Agricultural Research Service, Washington, DC.
Greer, K. R., Hawkins, W. W., and Catlin, J. E. (1968). Experimental studies of controlled reproduction in elk (Wapiti). The Journal of Wildlife Management 32, 368–376.
| Experimental studies of controlled reproduction in elk (Wapiti).Crossref | GoogleScholarGoogle Scholar |
Gross, J. A., and Knight, J. E. (2000). Elk presence inside various-sized cattle exclosures. Journal of Range Management 53, 287–290.
| Elk presence inside various-sized cattle exclosures.Crossref | GoogleScholarGoogle Scholar |
Haggerty, J., and Travis, W. R. (2006). Out of administrative control: absentee owners, resident elk and the shifting nature of wildlife management in southwestern Montana. Geoforum 37, 816–830.
| Out of administrative control: absentee owners, resident elk and the shifting nature of wildlife management in southwestern Montana.Crossref | GoogleScholarGoogle Scholar |
Hamir, A. N., Kunkle, R. A., Miller, J. M., Greenlee, J. J., and Richt, J. A. (2006). Experimental second passage of chronic wasting disease (CWDmule deer) agent to cattle. Journal of Comparative Pathology 134, 63–69.
| Experimental second passage of chronic wasting disease (CWDmule deer) agent to cattle.Crossref | GoogleScholarGoogle Scholar | 1:STN:280:DC%2BD28%2Fjs1eqsg%3D%3D&md5=8cf11ffca93d81373b6920c0a6404b93CAS | 16423572PubMed |
Hams, K., and Trindle, B. (2008). Annual performance report: 2007–08. W-15-R, Segment 64. Nebraska Game and Parks Commission, Wildlife Division, Lincoln, NE.
Hansen, I., and Bakken, M. (1999). Livestock-guarding dogs in Norway: Part I. Interactions. Journal of Range Management 52, 2–6.
| Livestock-guarding dogs in Norway: Part I. Interactions.Crossref | GoogleScholarGoogle Scholar |
Hansen, L., and Beringer, J. (1997). Managed hunts to control white-tailed deer populations on urban public areas in Missouri. Wildlife Society Bulletin 25, 484–487.
Hargreaves, S. K., Foggin, C. M., Anderson, E. C., Bastos, A. D. S., Thomson, G. R., Ferris, N. P., and Knowles, N. J. (2004). An investigation into the source and spread of foot and mouth disease virus from a wildlife conservancy in Zimbabwe. Revue Scientifique et Technique (International Office of Epizootics) 23, 783–790.
| 1:STN:280:DC%2BD2M3islSqsg%3D%3D&md5=67b3955f914d67909e0f8db39ff6f4acCAS | 15861873PubMed |
Harrington, J. L., and Conover, M. R. (2006). Characteristics of ungulate behavior and mortality associated with wire fences. Wildlife Society Bulletin 34, 1295–1305.
| Characteristics of ungulate behavior and mortality associated with wire fences.Crossref | GoogleScholarGoogle Scholar |
Harris, M. T., Palmer, W. L., and George, J. L. (1983). Preliminary screening of white-tailed deer repellents. The Journal of Wildlife Management 47, 516–519.
| Preliminary screening of white-tailed deer repellents.Crossref | GoogleScholarGoogle Scholar |
Hebblewhite, M., and Merrill, E. H. (2009). Trade-offs between predation risk and forage differ between migrant strategies in a migratory ungulate. Ecology 90, 3445–3454.
| Trade-offs between predation risk and forage differ between migrant strategies in a migratory ungulate.Crossref | GoogleScholarGoogle Scholar | 20120812PubMed |
Hebblewhite, M., and Pletscher, D. H. (2002). Effects of elk group size on predation by wolves. Canadian Journal of Zoology 80, 800–809.
| Effects of elk group size on predation by wolves.Crossref | GoogleScholarGoogle Scholar |
Hebblewhite, M., Pletscher, D. H., and Paquet, P. C. (2002). Elk population dynamics in areas with and without predation by recolonizing wolves in Banff National Park, Alberta. Canadian Journal of Zoology 80, 789–799.
| Elk population dynamics in areas with and without predation by recolonizing wolves in Banff National Park, Alberta.Crossref | GoogleScholarGoogle Scholar |
Hebblewhite, M., Merrill, E. H., Morgantini, L. E., White, C. A., Allen, J. R., Bruns, E., Thurston, L., and Hurd, T. E. (2006). Is the migratory behavior of montane elk herds in peril? The case of Alberta’s Ya Ha Tinda elk herd. Wildlife Society Bulletin 34, 1280–1294.
| Is the migratory behavior of montane elk herds in peril? The case of Alberta’s Ya Ha Tinda elk herd.Crossref | GoogleScholarGoogle Scholar |
Hegel, T. M., Gates, C. C., and Eslinger, D. (2009). The geography of conflict between elk and agricultural values in the Cypress Hills, Canada. Journal of Environmental Management 90, 222–235.
| The geography of conflict between elk and agricultural values in the Cypress Hills, Canada.Crossref | GoogleScholarGoogle Scholar | 18082311PubMed |
Henigman, J., Turner, J., and Swift, K. (2005). Roosevelt elk wildlife habitat decision aid. BC Journal of Ecosystems and Management 6, 51–53.
Herner-Thogmartin, J. (1999). Ecology of an introduced Rocky Mountain elk (Cervus elaphus) herd in Arkansas. M.Sc. Thesis, University of Arkansas, Fayetteville, AR.
Hines, A. M., Ezenwa, V. O., Cross, P., and Rogerson, J. D. (2007). Effects of supplemental feeding on gastrointestinal parasite infection in elk (Cervus elaphus): preliminary observations. Veterinary Parasitology 148, 350–355.
| Effects of supplemental feeding on gastrointestinal parasite infection in elk (Cervus elaphus): preliminary observations.Crossref | GoogleScholarGoogle Scholar | 17692464PubMed |
Hofmann, R. R. (1988). Anatomy of the gastro-intestinal tract. In ‘The Ruminant Animal: Digestive Physiology and Nutrition’. (Ed. D. C. Church.) pp. 14–43. (Prentice-Hall: Englewood Cliffs, NJ.)
Holdø, R. M., Dudley, J. P., and McDowell, L. R. (2002). Geophagy in the African elephant in relation to availability of dietary sodium. Journal of Mammalogy 83, 652–664.
| Geophagy in the African elephant in relation to availability of dietary sodium.Crossref | GoogleScholarGoogle Scholar |
Hone, J. (1990). Predator–prey theory and feral pig control, with emphasis on evaluation of shooting from a helicopter. Australian Wildlife Research 17, 123–130.
| Predator–prey theory and feral pig control, with emphasis on evaluation of shooting from a helicopter.Crossref | GoogleScholarGoogle Scholar |
Hygnstrom, S. E., and Craven, S. R. (1988). Electric fences and commercial repellents for reducing deer damage in cornfields. Wildlife Society Bulletin 16, 291–296.
Hygnstrom, S. E., Cover, M. A., Stillings, B. A., Crank, R. D., Fischer, J. W., Merchant, J. W., and Korte, S. P. (2005). Elk in Nebraska: opportunity or another private–public land conundrum. Proceedings of the Wildlife Damage Management Conference 11, 213–218.
Ishmael, W. E., and Rongstad, O. J. (1984). Economics of an urban deer-removal program. Wildlife Society Bulletin 12, 394–398.
Ishmael, W. E., Katsma, D. E., Isaac, A., and Bryant, B. K. (1995). Live-capture and translocation of suburban white-tailed deer in River Hills, Wisconsin. Proceedings of the Midwest Fish and Wildlife Conference 55, 87–96.
Jay, M. T., Cooley, M., Carychao, D., Wiscomb, G. W., Sweitzer, R. A., Crawford-Miksza, L., Farrar, J. A., Lau, D. K., O’Connell, J., Millington, A., Asmundson, R. V., Atwill, E. R., and Mandrell, R. E. (2007). Escherichia coli O157:H7 in feral swine near spinach fields and cattle, central California coast. Emerging Infectious Diseases 13, 1908–1911.
| 18258044PubMed |
Jenkins, K. J., and Starkey, E. E. (1993). Winter forages and diets of elk in old-growth and regenerating coniferous forests in western Washington. American Midland Naturalist 130, 299–313.
| Winter forages and diets of elk in old-growth and regenerating coniferous forests in western Washington.Crossref | GoogleScholarGoogle Scholar |
Johnson, T. L., and Monello, R. J. (2001). An adaptive approach to elk management in Rocky Mountain National Park. In ‘Crossing Boundaries in Park Management: Proceedings of the Conference on Research and Resource Management in Parks and on Public Lands’. (Ed. D. Harmon.) pp. 82–86. (The George Wright Society: Hancock, MI.)
Jones, J. M., and Witham, J. H. (1990). Post-translocation survival and movements of metropolitan white-tailed deer. Wildlife Society Bulletin 18, 434–441.
Jordan, A. J., and Workman, J. P. (1989). Economics and management of fee hunting for deer and elk in Utah. Wildlife Society Bulletin 17, 482–487.
Kahler, K. E. (1991). Responsible strategies for wildlife control. Golf Course Management June, 72–80.
Karhu, R. R., and Anderson, S. H. (2006). The effect of high-tensile electric fence designs on big-game and livestock movements. Wildlife Society Bulletin 34, 293–299.
| The effect of high-tensile electric fence designs on big-game and livestock movements.Crossref | GoogleScholarGoogle Scholar |
Kay, C. E. (1994). Aboriginal overkill – the role of Native Americans in structuring western ecosystems. Human Nature 5, 359–398.
| Aboriginal overkill – the role of Native Americans in structuring western ecosystems.Crossref | GoogleScholarGoogle Scholar |
Kay, C. E. (1997). Viewpoint: ungulate herbivory, willows, and political ecology in Yellowstone. Journal of Range Management 50, 139–145.
| Viewpoint: ungulate herbivory, willows, and political ecology in Yellowstone.Crossref | GoogleScholarGoogle Scholar |
Kessler, K. K. (1995). Financial compensation for wildlife damage: a review of programs in North America. Great Plains Wildlife Damage Control Workshop Proceedings 12, 90.
Killian, G., Eisemann, J., Wagner, D., Werner, J., Shaw, D., Engeman, R., and Miller, L. (2006). Safety and toxicity evaluation of GonaCon immunocontraceptive vaccine in white-tailed deer. Proceedings of the Vertebrate Pest Conference 22, 82–87.
Killian, G., Kreeger, T. J., Rhyan, J., Fagerstone, K., and Miller, L. (2009). Observations on the use of Gonacon™ in captive female elk (Cervus elaphus). Journal of Wildlife Diseases 45, 184–188.
| 19204347PubMed |
Kilpatrick, H. J., Spohr, S. M., and Chasko, G. G. (1997). A controlled deer hunt on a state-owned coastal reserve in Connecticut: controversies, strategies, and results. Wildlife Society Bulletin 25, 451–456.
Kilpatrick, H. J., Spohr, S. M., and Lima, K. K. (2001). Effects of population reduction on home ranges of female white-tailed deer at high densities. Canadian Journal of Zoology 79, 949–954.
| Effects of population reduction on home ranges of female white-tailed deer at high densities.Crossref | GoogleScholarGoogle Scholar |
Kilpatrick, A. M., Gillin, C. M., and Daszak, P. (2009). Wildlife–livestock conflict: the risk of pathogen transmission from bison to cattle outside Yellowstone National Park. Journal of Applied Ecology 46, 476–485.
| Wildlife–livestock conflict: the risk of pathogen transmission from bison to cattle outside Yellowstone National Park.Crossref | GoogleScholarGoogle Scholar |
Kimball, B. A., and Nolte, D. L. (2006). Development of a new deer repellent for the protection of forest resources. Western Journal of Applied Forestry 21, 108–111.
Kimball, B. A., Nolte, D. L., and Perry, K. B. (2005). Hydrolyzed casein reduces browsing of trees and shrubs by white-tailed deer. HortScience 40, 1810–1814.
| 1:CAS:528:DC%2BD2MXhtVyht7vN&md5=b965ed9f17d8dd140089bb7644c7d445CAS |
Kloppers, E. L., Clair, C. C., and Hurd, T. E. (2005). Predator-resembling aversive conditioning for managing habituated wildlife. Ecology and Society 10, 31.
Knight, R. R., and Mudge, M. R. (1967). Characteristics of some natural licks in the Sun River Area, Montana. The Journal of Wildlife Management 31, 293–299.
| Characteristics of some natural licks in the Sun River Area, Montana.Crossref | GoogleScholarGoogle Scholar | 1:CAS:528:DyaF2sXksFWlsLg%3D&md5=b6c18be7bbf4244fc4722c11aaf7f2b0CAS |
Knight, J. E., Swensson, E. J., and Sherwood, H. (1997). Elk use of modified fence-crossing designs. Wildlife Society Bulletin 25, 819–822.
Knipling, E. F. (1959). Sterile-male method of population control. Science 130, 902–904.
| Sterile-male method of population control.Crossref | GoogleScholarGoogle Scholar | 1:STN:280:DyaF3c7mt1Orsg%3D%3D&md5=4e5c723f777581543a4cceb48c1b5007CAS | 14410136PubMed |
Koehler, A. E., Marsh, R. E., and Salmon, T. P. (1990). Frightening methods and devices/stimuli to prevent mammal damage – A review. Proceedings of the Vertebrate Pest Conference 14, 168–173.
Larkin, J. L., Maehr, D. S., Cox, J. J., Bolin, D. C., and Wichrowski, M. W. (2003). Demographic characteristics of a reintroduced elk population in Kentucky. The Journal of Wildlife Management 67, 467–476.
| Demographic characteristics of a reintroduced elk population in Kentucky.Crossref | GoogleScholarGoogle Scholar |
Larkin, J. L., Cox, J. J., Wichrowski, M. J., Dzialak, M. R., and Maehr, D. S. (2004). Influences on release site fidelity of translocated elk. Restoration Ecology 12, 97–105.
| Influences on release site fidelity of translocated elk.Crossref | GoogleScholarGoogle Scholar |
Lauber, T. B., Brown, T. L., and Gore, M. L. (2004). Learning by doing: deer management in urban and suburban communities. Human Dimensions Research Unit Series No. 04-2. Cornell University, Ithaca, NY.
Lee, M. E., and Miller, R. (2003). Managing elk in the wildland–urban interface: attitudes of Flagstaff, Arizona, residents. Wildlife Society Bulletin 31, 185–191.
Lehnert, M. E., and Bissonette, J. A. (1997). Effectiveness of highway crosswalk structures at reducing deer–vehicle collisions. Wildlife Society Bulletin 25, 809–818.
Licht, D. S., Millspaugh, J. J., Kunkel, K. E., Kochanny, C. O., and Peterson, R. O. (2010). Using small populations of wolves for ecosystem restoration and stewardship. Bioscience 60, 147–153.
| Using small populations of wolves for ecosystem restoration and stewardship.Crossref | GoogleScholarGoogle Scholar |
Long, W. M. (1989). Habitat manipulations to prevent elk damage to private rangelands. Proceedings of the Great Plains Wildlife Damage Control Workshop 9, 101–103.
Lovaas, A. L. (1973). A cooperative elk trapping program in Wind Cave National Park. Wildlife Society Bulletin 1, 93–100.
Loveridge, A. J., Reynolds, J. C., and Milner-Gulland, E. J. (2006). Does sport hunting benefit conservation? In ‘Key Topics in Conservation Biology’. (Eds D. W. Macdonald and K. Service.) pp. 224–240. (Oxford University Press: Oxford, UK.)
Lubow, B. C., Singer, F. J., Johnson, T. L., and Bowden, D. C. (2002). Dynamics of interacting elk populations within and adjacent to Rocky Mountain National Park. The Journal of Wildlife Management 66, 757–775.
| Dynamics of interacting elk populations within and adjacent to Rocky Mountain National Park.Crossref | GoogleScholarGoogle Scholar |
MacLean, R. A., Mathews, N. E., Grove, D. M., Frank, E. S., and Paul-Murphy, J. (2006). Surgical technique for tubal ligation in white-tailed deer (Odocoileus virginianus). Journal of Zoo and Wildlife Medicine 37, 354–360.
| Surgical technique for tubal ligation in white-tailed deer (Odocoileus virginianus).Crossref | GoogleScholarGoogle Scholar | 17319135PubMed |
Maehr, D. S., Grimes, R., and Larkin, J. L. (1999). Initiating elk restoration: the Kentucky case study. Proceedings of the Annual Conference of the Southeastern Association of Game and Fish Agencies 53, 350–363.
Malcolm, K. D., Van Deelen, T. R., Drake, D., Kesler, D. J., and VerCauteren, K. C. (2010). Contraceptive efficacy of a novel intrauterine device (IUD) in white-tailed deer. Animal Reproduction Science 117, 261–265.
| Contraceptive efficacy of a novel intrauterine device (IUD) in white-tailed deer.Crossref | GoogleScholarGoogle Scholar | 19497690PubMed |
Mason, R. J. (1998). Mammal repellants: options and considerations for development. Proceedings of the Vertebrate Pest Conference 18, 325–329.
Mauldin, R. E., and Miller, L. A. (2007). Wildlife contraception: targeting the oocyte. In ‘Proceedings of an International Symposium on Managing Vertebrate Invasive Species’. (Eds G. W. Witmer, W. C. Pitt and K. A. Fagerstone.) pp. 434–444. (USDA/APHIS/WS National Wildlife Research Center: Fort Collins, CO.)
McCabe, R. E. (2002). Elk and Indians: then again. In ‘North American Elk: Ecology and Management’. (Eds D. E. Toweill and J. W. Thomas.) pp. 121–197. (Smithsonian Institute Press: Washington, DC.)
McCorquodale, S. M. (1987). Fall–Winter habitat use by elk in the shrub-steppe of Washington. Northwest Science 61, 171–173.
McCorquodale, S. M. (1991). Energetic considerations and habitat quality for elk in arid grasslands and coniferous forests. The Journal of Wildlife Management 55, 237–242.
| Energetic considerations and habitat quality for elk in arid grasslands and coniferous forests.Crossref | GoogleScholarGoogle Scholar |
McKenzie, J. A. (2001). The demographic and nutritional benefits of urban habitat use by elk. M.Sc. Thesis, University of Guelph, Ontario, Canada.
McKillop, I. G., and Sibly, R. M. (1988). Animal behaviour at electric fences and implications for management. Mammal Review 18, 91–103.
| Animal behaviour at electric fences and implications for management.Crossref | GoogleScholarGoogle Scholar |
Miller, L. A., Johns, B. E., Elias, D. J., and Killian, G. J. (1999). Oral vaccination of white-tailed deer using a recombinant Bacillus Calmette-Guerin vaccine expressing the Borrelia burgdorferi outer surface protein A: prospects for immunocontraception. American Journal of Reproductive Immunology 41, 279–285.
| 1:STN:280:DyaK1MzgtFansw%3D%3D&md5=f5518e416117bd427353503a0bc59f53CAS | 10374705PubMed |
Miller, R., Kaneene, J. B., Fitzgerald, S. D., and Schmitt, S. M. (2003). Evaluation of the influence of supplemental feeding of white-tailed deer (Odocoileus virginianus) on the prevalence of bovine tuberculosis in the Michigan wild deer population. Journal of Wildlife Diseases 39, 84–95.
| 12685071PubMed |
Miller, L. A., Gionfriddo, J. P., Fagerstone, K. A., Rhyan, J. C., and Killian, G. J. (2008). The single-shot GnRH immunocontraceptive vaccine (GonaCon) in white-tailed deer: comparison of several GnRH preparations. American Journal of Reproductive Immunology 60, 214–223.
| The single-shot GnRH immunocontraceptive vaccine (GonaCon) in white-tailed deer: comparison of several GnRH preparations.Crossref | GoogleScholarGoogle Scholar | 1:CAS:528:DC%2BD1cXhtFKrsbzP&md5=66115397852b0d6287e1000b36f7c66cCAS | 18782282PubMed |
Miller, L. A., Fagerstone, K. A., Wagner, D. C., and Killian, G. J. (2009). Factors contributing to the success of a singl-shot, multiyear PZP immunocontraceptive vaccine for white-tailed deer. Human–Wildlife Conflicts 3, 103–115.
Morrison, J. R., de Vergie, W. J., Alldredge, A. W., Byrne, A. E., and Andree, W. W. (1995). The effects of ski area expansion on elk. Wildlife Society Bulletin 23, 481–489.
Mould, E. D., and Robbins, C. T. (1981). Nitrogen metabolism in elk. The Journal of Wildlife Management 45, 323–334.
| Nitrogen metabolism in elk.Crossref | GoogleScholarGoogle Scholar |
Mould, E. D., and Robbins, C. T. (1982). Digestive capabilities in elk compared to white-tailed deer. The Journal of Wildlife Management 46, 22–29.
| Digestive capabilities in elk compared to white-tailed deer.Crossref | GoogleScholarGoogle Scholar |
Muller-Schwarze, D. (1994). The senses of deer. In ‘The Wildlife Series: Deer’. (Eds D. Gerlach, S. Atwater and J. Schnell.) pp. 58–665. (Stackpole Books: Mechanicsburg, PA.)
Neff, M. W. (2007). Scenario planning for wildlife management: a case study of the National Elk Refuge, Jackson, Wyoming. Human Dimensions of Wildlife 12, 219–226.
| Scenario planning for wildlife management: a case study of the National Elk Refuge, Jackson, Wyoming.Crossref | GoogleScholarGoogle Scholar |
Nolte, D. L. (1998). Efficacy of selected repellents to deter deer browsing on conifer seedlings. International Biodeterioration & Biodegradation 42, 101–107.
| Efficacy of selected repellents to deter deer browsing on conifer seedlings.Crossref | GoogleScholarGoogle Scholar | 1:CAS:528:DyaK1cXnvVOntLY%3D&md5=03dd25d93a613d104b46cd9dc7468292CAS |
Nolte, D. (1999). Behavioral approaches for limiting depredation by wild ungulates. In ‘Grazing Behavior of Livestock and Wildlife’. (Eds K. L. Launchbaugh, K. D. Sanders and J. C. Mosley.) pp. 60–69. Idaho Forest, Wildlife & Range Experimental Station Bulletin #70. (University of Idaho: Moscow, ID.)
Nolte, D. L., Farley, J. P., and Holbrook, S. (1995). Effectiveness of BGR-P and garlic in inhibiting browsing of western redcedar by black-tailed deer. Tree Planters’ Notes 46, 23–27.
Nolte, D. L., Shipley, L. A., and Wagner, K. K. (2001). Efficacy of Wolfin to repel black-tailed deer. Western Journal of Applied Forestry 16, 182–186.
Nolte, D. L., VerCauteren, K. C., Perry, K. R., and Adams, S. E. (2003). Training deer to avoid sites through negative reinforcement. Proceedings of the Wildlife Damage Management Conference 10, 95–104.
Nugent, G., and Choquenot, D. (2004). Comparing cost-effectiveness of commercial harvesting, state-funded culling, and recreational deer hunting in New Zealand. Wildlife Society Bulletin 32, 481–492.
| Comparing cost-effectiveness of commercial harvesting, state-funded culling, and recreational deer hunting in New Zealand.Crossref | GoogleScholarGoogle Scholar |
Nugent, G., Fraser, K. W., Asher, G. W., and Tustin, K. G. (2001). Advances in New Zealand mammalogy 1990–2000: deer. Journal of the Royal Society of New Zealand 31, 263–298.
| Advances in New Zealand mammalogy 1990–2000: deer.Crossref | GoogleScholarGoogle Scholar |
O’Bryan, M. K., and McCullough, D. R. (1985). Survival of black-tailed deer following relocation in California. The Journal of Wildlife Management 49, 115–119.
| Survival of black-tailed deer following relocation in California.Crossref | GoogleScholarGoogle Scholar |
O’Gara, B. W., and Dundas, R. G. (2002). Distribution: past and present. In ‘North American Elk: Ecology and Management’. (Eds D. E. Toweill and J. W. Thomas.) pp. 67–119. (Smithsonian Institute Press: Washington, DC.)
Oldenburg, L. E. (1984). Idaho elk population status/depredation problems. In ‘Western States Elk Workshop’. (Ed. R. W. Nelson.) pp. 75–79. (Western Association of Fish and Wildlife Agencies and Alberta Fish and Wildlife Division: Edmonton, Alberta.)
Palmer, W. L., Wingard, R. G., and George, J. L. (1983). Evaluation of white-tailed deer repellents. Wildlife Society Bulletin 11, 164–166.
Parkes, J. P., Nugent, G., and Warburton, B. (1996). Commercial exploitation as a pest control tool for introduced mammals in New Zealand. Wildlife Biology 2, 171–177.
Peek, J. M., Schmidt, K. T., Dorrance, M. J., and Smith, B. L. (2002). Supplemental feeding and farming of elk. In ‘North American Elk: Ecology and Management’. (Eds D. E. Toweill and J. W. Thomas.) pp. 617–647. (Smithsonian Institute Press: Washington, DC.)
Peinetti, H. R., Kalkhan, M. A., and Coughenour, M. B. (2002). Long-term changes in willow spatial distribution on the elk winter range of Rocky Mountain National Park (USA). Landscape Ecology 17, 341–354.
| Long-term changes in willow spatial distribution on the elk winter range of Rocky Mountain National Park (USA).Crossref | GoogleScholarGoogle Scholar |
Peterson, M. N., Lopez, R. R., Silvy, N. J., Owen, C. B., Frank, P. A., and Braden, A. W. (2003). Evaluation of deer-exclusion grates in urban areas. Wildlife Society Bulletin 31, 1198–1204.
Phillips, G. E., and Alldregde, A. W. (2000). Reproductive success of elk following disturbance by humans during calving season. The Journal of Wildlife Management 64, 521–530.
| Reproductive success of elk following disturbance by humans during calving season.Crossref | GoogleScholarGoogle Scholar |
Plumb, G. E., White, P. J., Coughenour, M. B., and Wallen, R. L. (2009). Carrying capacity, migration, and dispersal in Yellowstone bison. Biological Conservation 142, 2377–2387.
| Carrying capacity, migration, and dispersal in Yellowstone bison.Crossref | GoogleScholarGoogle Scholar |
Porter, W. F. (1983). A baited electric fence for controlling deer damage to orchard seedlings. Wildlife Society Bulletin 11, 325–327.
Putman, R. J., and Staines, B. W. (2004). Supplementary winter feeding of wild red deer Cervus elaphus in Europe and North America: justifications, feeding practice and effectiveness. Mammal Review 34, 285–306.
| Supplementary winter feeding of wild red deer Cervus elaphus in Europe and North America: justifications, feeding practice and effectiveness.Crossref | GoogleScholarGoogle Scholar |
Ramsey, C. W. (1968). A drop-net deer trap. The Journal of Wildlife Management 32, 187–190.
| A drop-net deer trap.Crossref | GoogleScholarGoogle Scholar |
Ramsey, K. J., and Krueger, W. C. (1986). Grass–legume seeding to improve winter forage for Roosevelt elk: a literature review. Agricultural Experiment Station Special Report 763. Oregon State University, Corvallis, OR.
Relyea, R. A., Lawrence, R. K., and Demarais, S. (2000). Home range of desert mule deer: testing the body-size and habitat-productivity hypotheses. The Journal of Wildlife Management 64, 146–153.
| Home range of desert mule deer: testing the body-size and habitat-productivity hypotheses.Crossref | GoogleScholarGoogle Scholar |
Rigg, R. (2001). Livestock guarding dogs: their current use world wide. Canid Specialist Group Occassional Paper No. 1. Available at http://www.canids.org/occasionalpapers/ [accessed 17 January 2010].
Ripple, W. J., and Beschta, R. L. (2003). Wolf reintroduction, predation risk and cottonwood recovery in Yellowstone National Park. Forest Ecology and Management 184, 299–313.
| Wolf reintroduction, predation risk and cottonwood recovery in Yellowstone National Park.Crossref | GoogleScholarGoogle Scholar |
Roper, R. B., and Hill, E. P. (1985). An evaluation of visual and auditory electronic devices to repel deer. Proceedings of the Eastern Wildlife Damage Control Conference 2, 186–191.
Rosatte, R., Hamr, J., Young, J., Filion, I., and Smith, H. (2007). The restoration of elk (Cervus elaphus) in Ontario, Canada: 1998–2005. Restoration Ecology 15, 34–43.
| The restoration of elk (Cervus elaphus) in Ontario, Canada: 1998–2005.Crossref | GoogleScholarGoogle Scholar |
Rowland, M. M., Wisdom, M. J., Johnson, B. K., and Kie, J. G. (2000). Elk distribution and modeling in relation to roads. The Journal of Wildlife Management 64, 672–684.
| Elk distribution and modeling in relation to roads.Crossref | GoogleScholarGoogle Scholar |
Rubin, E. S., Boyce, W. M., Stermer, C. J., and Torres, S. G. (2002). Bighorn sheep habitat use and selection near an urban environment. Biological Conservation 104, 251–263.
| Bighorn sheep habitat use and selection near an urban environment.Crossref | GoogleScholarGoogle Scholar |
Rutberg, A. T., Naugle, R. E., Thiele, L. A., and Liu, I. K. M. (2004). Effects of immunocontraception on a suburban population of white-tailed deer Odocoileus virginianus. Biological Conservation 116, 243–250.
| Effects of immunocontraception on a suburban population of white-tailed deer Odocoileus virginianus.Crossref | GoogleScholarGoogle Scholar |
Sayre, R. W., and Richmond, M. E. (1992). Evaluation of a new deer repellent on Japanese yews at suburban homesites. Proceedings of the Eastern Wildlife Damage Control Conference 5, 38–93.
Schemnitz, S. D. (1994). Capturing and handling wild animals. In ‘Research and Management Techniques for Wildlife and Habitats’. (Ed. T. A. Bookhout.) pp. 106–124. (The Wildlife Society: Bethesda, MD.)
Schneidmiller, J. F. (1987). Fencing methods to control big game damage to stored crops in Wyoming. Proceedings of the Great Plains Wildlife Damage Control Workshop 8, 217–221.
Schoenecker, K. A., Singer, F. J., Zeigenfuss, L. C., Binkley, D., and Menezes, R. S. C. (2004). Effects of elk herbivory on vegetation and nitrogen processes. The Journal of Wildlife Management 68, 837–849.
| Effects of elk herbivory on vegetation and nitrogen processes.Crossref | GoogleScholarGoogle Scholar |
Schultz, R. D., and Bailey, J. A. (1978). Responses of National Park elk to human activity. The Journal of Wildlife Management 42, 91–100.
| Responses of National Park elk to human activity.Crossref | GoogleScholarGoogle Scholar |
Scott, M. D. (1992). Buck-and-pole fence crossings by 4 ungulate species. Wildlife Society Bulletin 20, 204–210.
Seamans, T. W., and Helon, D. A. (2008). Comparison of electrified mats and cattle guards to control white-tailed deer (Odocoileus virginianus) access through fences. Proceedings of the Vertebrate Pest Conference 23, 206–209.
Seamans, T. W., Blackwell, B. F., and Cepek, J. D. (2002). Coyote hair as an area repellent for white-tailed deer. International Journal of Pest Management 48, 301–306.
| Coyote hair as an area repellent for white-tailed deer.Crossref | GoogleScholarGoogle Scholar |
Seward, N. W., Phillips, G. E., Duquette, J. F., and Vercauteren, K. C. (2007). A frightening device for deterring deer use of cattle feeders. The Journal of Wildlife Management 71, 271–276.
| A frightening device for deterring deer use of cattle feeders.Crossref | GoogleScholarGoogle Scholar |
Shively, K. J., Alldregde, A. W., and Phillips, G. E. (2005). Elk reproductive response to removal of calving season disturbance by humans. The Journal of Wildlife Management 69, 1073–1080.
| Elk reproductive response to removal of calving season disturbance by humans.Crossref | GoogleScholarGoogle Scholar |
Shivik, J. A., and Martin, D. J. (2000). Aversive and disruptive stimulus applications for managing predation. Proceedings of the Wildlife Damage Management Conference 9, 111–119.
Short, H. L., Dietz, D. R., and Remmenga, E. E. (1966). Selected nutrients in mule deer browse plants. Ecology 47, 222–229.
| Selected nutrients in mule deer browse plants.Crossref | GoogleScholarGoogle Scholar |
Skovlin, J. M., Edgerton, P. J., and McConnell, B. R. (1983). Elk use of winter range as affected by cattle grazing, fertilizing, and burning in southeastern Washington. Journal of Range Management 36, 184–189.
| Elk use of winter range as affected by cattle grazing, fertilizing, and burning in southeastern Washington.Crossref | GoogleScholarGoogle Scholar |
Smith, B. L. (2001). Winter feeding of elk in western North America. The Journal of Wildlife Management 65, 173–190.
| Winter feeding of elk in western North America.Crossref | GoogleScholarGoogle Scholar |
Smith, R. H., Neff, D. J., and Woolsey, N. G. (1986). Pronghorn response to coyote control: a benefit : cost analysis. Wildlife Society Bulletin 14, 226–231.
Spalinger, D. E., Cooper, S. M., Martin, D. J., and Shipley, L. A. (1997). Is social learning an important influence on foraging behavior in white-tailed deer? The Journal of Wildlife Management 61, 611–621.
| Is social learning an important influence on foraging behavior in white-tailed deer?Crossref | GoogleScholarGoogle Scholar |
St Clair, C. C., and Forrest, A. (2009). Impacts of vehicle traffic on the distribution and behaviour of rutting elk, Cervus elaphus. Behaviour 146, 393–413.
| Impacts of vehicle traffic on the distribution and behaviour of rutting elk, Cervus elaphus.Crossref | GoogleScholarGoogle Scholar |
Stewart, S. C., Fix, P. J., and Manfredo, M. J. (2004). Public perceptions of elk and vegetation management in Rocky Mountain National Park, Colorado. Rocky Mountain National Park Project Report No. 56. Colorado State University, Fort Collins, CO.
Stockstad, D. S., Morris, M. S., and Lory, E. C. (1953). Chemical characteristics of natural licks used by big game animals in western Montana. Transactions of the North American Wildlife Conference 18, 247–257.
Storlie, J. T. (2006). Movements and habitat use of female Roosevelt elk in relation to human disturbance on the Hoko and Dickey Game Management Units, Washington. M.Sc. Thesis, Humbolt State University, Arcata, CA.
Stout, G. G., Lowry, F. C., and Carlisle, F. (1972). The status of elk transplants in eastern Oklahoma. Proceedings of the Annual Conference of the Southeastern Association of Game and Fish Agencies 26, 202–203.
Suzuki, K., Suzuki, H., Binkley, D., and Stohlgren, T. J. (1999). Aspen regeneration in the Colorado Front Range: differences at local and landscape scales. Landscape Ecology 14, 231–237.
| Aspen regeneration in the Colorado Front Range: differences at local and landscape scales.Crossref | GoogleScholarGoogle Scholar |
Swihart, R. K., and Conover, M. R. (1990). Reducing deer damage to yews and apple trees: testing Big Game Repellent®, RoPel® and soap as repellents. Wildlife Society Bulletin 18, 156–162.
Swihart, R. K., Pignatello, V., Mattina, M. J. I., and Tennessen, M. (1991). Aversive responses of white-tailed deer, Odocoileus virginianus, to predator urines. Journal of Chemical Ecology 17, 767–777.
| Aversive responses of white-tailed deer, Odocoileus virginianus, to predator urines.Crossref | GoogleScholarGoogle Scholar |
Taylor, T. S., Loewenstein, E. F., and Chappelka, A. H. (2006). Effect of animal browse protection and fertilizer application on the establishment of planted Nuttall oak seedlings. New Forests 32, 133–143.
| Effect of animal browse protection and fertilizer application on the establishment of planted Nuttall oak seedlings.Crossref | GoogleScholarGoogle Scholar |
Thompson, M. J., and Henderson, R. E. (1998). Elk habituation as a credibility challenge for wildlife professionals. Wildlife Society Bulletin 26, 477–483.
Thompson, M. J., Henderson, R. E., Lemke, T. O., and Sterling, B. A. (1989). Evaluation of a collapsible clover trap for elk. Wildlife Society Bulletin 17, 287–290.
Trombulak, S. C., and Frissell, C. A. (2000). Review of ecological effects of roads on terrestrial and aquatic communities. Conservation Biology 14, 18–30.
| Review of ecological effects of roads on terrestrial and aquatic communities.Crossref | GoogleScholarGoogle Scholar |
Tufto, J., Andersen, R., and Linnell, J. (1996). Habitat use and ecological correlates of home range size in a small cervid: the roe deer. Journal of Animal Ecology 65, 715–724.
| Habitat use and ecological correlates of home range size in a small cervid: the roe deer.Crossref | GoogleScholarGoogle Scholar |
Van Deelen, T. R. (2003). Chronic wasting disease and the science in support of the ban on baiting and feeding deer. Available at http://www.dnr.state.wi.us/org/land/wildlife/Whealth/issues/Cwd/doc/cwdscsu.pdf [accessed 12 April 2007].
Van Deelen, T. R., McKinney, L. B., Joselyn, M. G., and Buhnerkempe, J. E. (1997). Can we restore elk to southern Illinois? The use of existing digital land-cover data to evaluate potential habitat. Wildlife Society Bulletin 25, 886–894.
Van Tassell, L. W., Phillips, C., and Yang, B. (1999). Depredation claim settlements in Wyoming. Wildlife Society Bulletin 27, 479–487.
VerCauteren, K. C., and Pipas, M. J. (2003). A review of color vision in white-tailed deer. Wildlife Society Bulletin 31, 684–691.
VerCauteren, K. C., Beringer, J., and Hygnstrom, S. E. (1997). Use of netted-cage traps in population management and research of urban white-tailed deer. Proceedings of the Great Plains Wildlife Damage Control Workshop 13, 138–142.
VerCauteren, K. C., Hygnstrom, S. E., Pipas, M. J., Fioranelli, P. B., Werner, S. J., and Blackwell, B. F. (2003). Red lasers are ineffective for dispersing deer at night. Wildlife Society Bulletin 31, 247–252.
VerCauteren, K. C., Seward, N. W., Hirchert, D. L., Jones, M. L., and Beckerman, S. F. (2005a). Dogs for reducing wildlife damage to organic crops: a case study. Proceedings of the Wildlife Damage Management Conference 11, 286–293.
VerCauteren, K. C., Shivik, J. A., and Lavelle, M. J. (2005b). Efficacy of an animal-activated frightening device on urban elk and mule deer. Wildlife Society Bulletin 33, 1282–1287.
| Efficacy of an animal-activated frightening device on urban elk and mule deer.Crossref | GoogleScholarGoogle Scholar |
VerCauteren, K. C., Gilsdorf, J. M., Hygnstrom, S. E., Fioranelli, P. B., Wilson, J. A., and Barras, S. (2006a). Green and blue lasers are ineffective for dispersing deer at night. Wildlife Society Bulletin 34, 371–374.
| Green and blue lasers are ineffective for dispersing deer at night.Crossref | GoogleScholarGoogle Scholar |
VerCauteren, K. C., Lavelle, M. J., and Hygnstrom, S. E. (2006b). Fences and deer-damage management: a review of designs and efficacy. Wildlife Society Bulletin 34, 191–200.
| Fences and deer-damage management: a review of designs and efficacy.Crossref | GoogleScholarGoogle Scholar |
VerCauteren, K. C., Lavelle, M. J., and Hygnstrom, S. E. (2006c). A simulation model for determining cost-effectiveness of fences for reducing deer damage. Wildlife Society Bulletin 34, 16–22.
| A simulation model for determining cost-effectiveness of fences for reducing deer damage.Crossref | GoogleScholarGoogle Scholar |
VerCauteren, K. C., Lavelle, M. J., Seward, N. W., Fischer, J. W., and Phillips, G. E. (2007a). Fence-line contact between wild and farmed cervids in Colorado: potential for disease transmission. The Journal of Wildlife Management 71, 1594–1602.
| Fence-line contact between wild and farmed cervids in Colorado: potential for disease transmission.Crossref | GoogleScholarGoogle Scholar |
VerCauteren, K. C., Seward, N. W., Lavelle, M. J., Fischer, J. W., and Phillips, G. E. (2007b). A fence design for excluding elk without impeding other wildlife. Rangeland Ecology and Management 60, 529–532.
| A fence design for excluding elk without impeding other wildlife.Crossref | GoogleScholarGoogle Scholar |
VerCauteren, K. C., Lavelle, M. J., and Phillips, G. E. (2008). Livestock protection dogs for deterring deer from cattle and feed. The Journal of Wildlife Management 72, 1443–1448.
| Livestock protection dogs for deterring deer from cattle and feed.Crossref | GoogleScholarGoogle Scholar |
VerCauteren, K. C., Seward, N. W., Lavelle, M. J., Fischer, J. W., and Phillips, G. E. (2009). Deer guards and bump gates for excluding white-tailed deer from fenced resources. Human–Wildlife Conflicts 3, 145–153.
Vieira, M. E. P., Conner, M. M., White, G. C., and Freddy, D. J. (2003). Effects of archery hunter numbers and opening dates on elk movement. The Journal of Wildlife Management 67, 717–728.
| Effects of archery hunter numbers and opening dates on elk movement.Crossref | GoogleScholarGoogle Scholar |
Wagner, K. K., Schmidt, R. H., and Conover, M. R. (1997). Compensation programs for wildlife damage in North America. Wildlife Society Bulletin 25, 312–319.
Walter, W. D., and Leslie, D. M. (2002). Harvest strategies and numbers of elk (Cervus elaphus) in Oklahoma, 1987–2001. Proceedings of the Oklahoma Academy of Science 82, 89–94.
Walter, W. D., Pekins, P. J., Rutberg, A. T., and Kilpatrick, H. J. (2002). Evaluation of immunocontraception in suburban white-tailed deer. Wildlife Society Bulletin 30, 186–192.
Walter, W. D., Leslie, D. M., and Jenks, J. A. (2006). Response of Rocky Mountain elk (Cervus elaphus) to wind-power development. American Midland Naturalist 156, 363–375.
| Response of Rocky Mountain elk (Cervus elaphus) to wind-power development.Crossref | GoogleScholarGoogle Scholar |
Ward, A. L. (1982). Mule deer behavior in relation to fencing and underpasses on Interstate 80 in Wyoming. Transportation Research Record 859, 8–13.
Washington Department of Fish and Wildlife (2009). 2009 game status and trend report. Wildlife Program, Washington Department of Fish and Wildlife, Olympia, WA.
White, C. A., Olmsted, C. E., and Kay, C. E. (1998). Aspen, elk and fire in the Rocky Mountain National Parks of North America. Wildlife Society Bulletin 26, 449–462.
Whyte, I. J., and Joubert, S. C. J. (1988). Blue wildebeest population trends in the Kruger National Park and the effects of fencing. South African Journal of Wildlife Research 18, 78–87.
Williams, T. R. (1962). The significance of salt and natural licks in elk management. M.Sc. Thesis, University of Idaho, Moscow, ID.
Wilson, C. J., Britton, A. M., and Symes, R. G. (2009). An assessment of agricultural damage caused by red deer (Cervus elaphus L.) and fallow deer (Dama dama L.) in southwest England. Wildlife Biology in Practice 5, 104–114.
| An assessment of agricultural damage caused by red deer (Cervus elaphus L.) and fallow deer (Dama dama L.) in southwest England.Crossref | GoogleScholarGoogle Scholar |
Witham, J. (1996). Teaching elk to jump. Wildlife Views November, 17–19.
Witmer, G. W. (1998). When deer are too dear and elk are too elegant. The Probe 189, 1–5.
Witmer, G. W., and Cogan, R. (1989). Elk and crop damage in Pennsylvania. Proceedings of the Eastern Wildlife Damage Control Conference 4, 220–224.
Woodruff, R. A., and Green, J. S. (1995). Livestock herding dogs: a unigue application for wildlife damage management. Proceedings of the Great Plains Wildlife Damage Control Workshop 12, 43–45.
